J. exp. Biol. 157, 75-86 (1991) 75
Printed in Great Britain © The Company of Biologists Limited 1991
THE INFLUENCE OF HYPOXIA ON THE PREFERRED
TEMPERATURE OF RAINBOW TROUT
ONCORHYNCHUS MYKISS
BY H. SCHURMANN
1
, J. F. STEFFENSEN
2
* AND J. P. LOMHOLT
1
1
Department of Zoophysiology, University of Aarhus, DK-8000 Aarhus and
2
Marine Biological Laboratory, University of Copenhagen, DK-3000 Helsing0r,
Denmark
Accepted 17 December 1990
Summary
The behavioural thermoregulation of rainbow trout Oncorhynchus mykiss was
investigated in a shuttlebox at normoxia and at three levels of hypoxia: 13.3, 9.3
and 5.3 kPa. The final preferred temperatures at normoxia, and at 13.3, 9.3 and
5.3 kPa, were 16.1, 14.9, 15.0 and 12.7°C, respectively. A decreased rate of
metabolism and an increased blood oxygen-affinity are among the physiological
advantages of selecting a lower temperature during hypoxia. It is suggested that
catecholamines may play a part in controlling temperature selection during
hypoxia. In natural environments, this behaviour may result in habitat shifts of
fish living in heterothermal environments with changing oxygen tensions.
Introduction
Fish placed in a temperature gradient will initially select temperatures that
depend on the acclimation temperature. The selected temperature during the first
2 h in the gradient is termed the acute preferred temperature. If left in the gradient
for a longer period, usually 24 h (Reynolds and Casterlin, 1980), the fish will select
a species-specific preferred temperature. This temperature is termed the final
preferendum and is defined by Fry (1947) as 'the temperature at which the
individuals will ultimately congregate regardless of their thermal experience
before being placed in the gradient, and at which acclimation temperature and
acute preferred temperatures are equal'. Usually the selected temperature consists
of a range of temperatures bounded by an upper and lower avoidance tempera-
ture, so the final preferendum is actually a statistical measure of central tendency
in the temperature distribution (Reynolds and Casterlin, 1979).
Recently, the acute preferred temperature for fish (Bryan et al. 1984) has been
shown to decrease as a result of exposure to hypoxia. Similar results were obtained
in reptiles (Hicks and Wood, 1985), salamanders and crayfish (Dupre and Wood,
*To whom reprint requests should be addressed.
Key words: hypoxia, rainbow trout, preferred temperature, Oncorhynchus mykiss.