Russian Geology Geologiya
and Geophysics i Geofizika
Vol. 44, No. 7, pp. 664-675, 2003 UDC 591:551.762(265)
PALEOBIOCHORES OF JURASSIC BOREAL BASINS
V.A. Zakharov, S.V. Meledina*, and B.N. Shurygin*
Geological Institute of the RAS, 7 Pyzhevsky per., Moscow, 109017, Russia
* Institute of Petroleum Geology, Siberian Branch of the RAS,
3 prosp. Akad. Koptuyga, Novosibirsk, 630090, Russia
Paleobiogeographical zonation is usually based on a biochorological, or biogeographical,
method, which is reduced to mapping of ranges of individual taxa (families, genera, and
species), followed by their analysis. No unified nomenclature of biochores is available,
because there are no approved rules for paleobiogeographical zonation. The problem of
relationship between the rank of biochore and the rank of its indicator taxon has not been
solved yet. A common rule is that the rank of a biochore depends on the rank of taxonomic
distinctions. The rank Realm corresponds to the ranks Family and Superfamily of endemic
taxa in Russian systematics; the criterion for recognizing Subrealms is mainly distinctions
of generic level. Realms may be subdivided into Provinces, spatially separated communities
whose specific character is determined by species (subgeneric, generic) distinctions. To solve
the problem of the rank of biochores, it is necessary to analyze the history of the biota for
a long time. It seems reasonable to submit the following hierarchical series of the main
biogeographical categories: Superrealm, Realm, Province, with intermediate categories:
Subrealm and Subprovince; the term region is for informal use.
The largest biogeographical unit is Superrealm. Only synthesis of areal data on nektonic,
benthic, and planktonic groups permits recognition of general biogeographical regularities
and, therefore, makes the paleobiogeographical division more objective. First of all, this is
true for Superrealms and Realms. Recognition of a biochore should rely first of all on a
certain chorotype. Chorotype can be that part of a biochore where its taxonomic originality
is highlighted. The time of existence of the chorotype (chronotype) must also be provided.
The rule of priority name is required.
Biochores are to be accompanied with certain information. The chorotype designation should
be supplemented by the name of the first author(s) who proposed the biochore, initial time
for which it was established, its territory; paleontological description; additions introduced
into these attributes in subsequent publications. The present paper reports examples of
description of some chorotypes.
Mesozoic, Arctic, paleobiogeography, hierarchy of zoochores, principles of zonation
INTRODUCTION
At present, there are no universally accepted hierarchy and nomenclature of the paleobiochores* from Jurassic
Boreal basins. Paleobiogeographical zonation is usually carried out by experts specialized in different groups of
sea organisms independently of each other. The rank of biochores is treated in a subjective way, and their boundaries
do not always coincide with the boundaries earlier proposed by other authors. A frequent situation is that in drawing
the boundaries of a biochore, investigators encounter the problem of invariancy. A personal approach to solving
the problem is common since there are neither distinct boundaries between adjacent biochores nor strong rules of
their naming. In some cases, taxonomically and spatially close biochores are ranked and named differently.
©2003 UIGGM, SIBERIAN BRANCH OF THE RAS
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* The Friends of Paleobiogeography adopted the term (paleo)biochoreme, as proposed by M. Macenido, instead of (paleo)biochore.