Behavioural Processes 68 (2005) 179–184
Short report
Left hemispheric advantage for numerical
abilities in the bottlenose dolphin
Annette Kilian
a,b,*
, Lorenzo von Fersen
b
, Onur G ¨ unt¨ urk¨ un
a
a
Department Biopsychology, Inst. Cogn. Neurosci., Faculty of Psychology, Ruhr-University Bochum, 44780 Bochum, Germany
b
Tiergarten N¨ urnberg, Am Tiergarten 30, 90480 N¨ urnberg, Germany
Received 8 September 2004; received in revised form 23 October 2004; accepted 7 November 2004
Abstract
In a two-choice discrimination paradigm, a bottlenose dolphin discriminated relational dimensions between visual numerosity
stimuli under monocular viewing conditions. After prior binocular acquisition of the task, two monocular test series with different
number stimuli were conducted. In accordance with recent studies on visual lateralization in the bottlenose dolphin, our results
revealed an overall advantage of the right visual field. Due to the complete decussation of the optic nerve fibers, this suggests a
specialization of the left hemisphere for analysing relational features between stimuli as required in tests for numerical abilities.
These processes are typically right hemisphere-based in other mammals (including humans) and birds. The present data provide
further evidence for a general right visual field advantage in bottlenose dolphins for visual information processing. It is thus
assumed that dolphins possess a unique functional architecture of their cerebral asymmetries.
© 2004 Elsevier B.V. All rights reserved.
Keywords: Bottlenose dolphin; Hemispheric specialization; Monocular vision; Numerical ability
1. Introduction
Over the last decades, a large body of experimental
studies has accumulated, demonstrating that functional
asymmetries are not unique to humans, but represent
a characteristic that many vertebrate species have in
common (Rogers and Andrew, 2002). This growing
*
Corresponding author. Tel.: +49 9123 963050;
fax: +49 9123 963049.
E-mail address: akilian@marine-mammal-cognition.de
(A. Kilian).
evidence for perceptual and cognitive lateralizations in
animals also provides the basis for understanding the
biological foundations of hemispheric specializations.
So far, only few studies have focused on lateralizations
in marine mammals like dolphins. Apart from findings
on lateralized motor functions in the bottlenose dol-
phin (Norris and Dohl, 1980; Marino and Stowe, 1997),
first investigations on cerebral functional asymmetries
were carried out by von Fersen et al. (2000) and Kilian
et al. (2000) demonstrating a right eye dominance in
two different visual tasks for their subjects. Due to the
complete decussation of the optic nerves in dolphins
0376-6357/$ – see front matter © 2004 Elsevier B.V. All rights reserved.
doi:10.1016/j.beproc.2004.11.003