Plant Herbivore Interactions
at the Forest Edge
Rainer Wirth ( ), Sebastian T. Meyer, Inara R. Leal,
and Marcelo Tabarelli
Contents
1 Introduction .......................................................................................................................... 424
2 What Edges Induce in Forest Remnants............................................................................... 425
3 Plant-Herbivore Interactions at the Edge: Patterns and Processes ....................................... 427
3.1 Patterns ........................................................................................................................ 427
3.2 Processes ..................................................................................................................... 432
4 Impact of Herbivory on the Forest Edge .............................................................................. 436
4.1 Abiotic Conditions ...................................................................................................... 437
4.2 Plant Community......................................................................................................... 438
4.3 Higher Trophic Levels ................................................................................................. 440
5 Conclusions and Outlook ..................................................................................................... 441
References .................................................................................................................................. 442
Abstract An ever-increasing proportion of the global forested landscape is in
close proximity to edges and edge effects have been shown to represent key forces
affecting both organisms and ecological processes. Despite increasing recognition
of edge effects on species interactions, a systematic review devoted to plant-
herbivore interactions along forest edges has not yet been performed. Here we syn-
thesize published research attempting to detect patterns of herbivore densities and
herbivory at forest edges, identify the underlying mechanisms generating these
patterns, and explore their potential impacts for the forest edge as an ecosystem.
Key conclusions are that herbivores, especially generalists, profoundly benefit
from forest edges, often due to favourable microenvironmental conditions, an
edge-induced increase in food quantity/quality, and (less well documented)
disrupted top-down regulation of herbivores. Finally, we present evidence and
causal explanations that edge-associated herbivores, via a range of direct and
indirect impacts, may alter species interactions, delay successional processes at
the edge, and amplify the often human-induced changes on forest biota.
Rainer Wirth
Plant Ecology & Systematics, University of Kaiserslautern, PO Box 3049, D-67663
Kaiserslautern, Germany
e-mail: wirth@rhrk.uni-kl.de
U. Lüttge et al. (eds.), Progress in Botany 69. 423
© Springer-Verlag Berlin Heidelberg 2008