Current Neurovascular Research, 2009, 6, 259-266 259
1567-2026/09 $55.00+.00 © 2009 Bentham Science Publishers Ltd.
Chronic Methylphenidate-Effects Over Circadian Cycle of Young and
Adult Rats Submitted to Open-Field and Object Recognition Tests
Karin M. Gomes
1
, Renan P. Souza
2
, Samira S. Valvassori
1
, Gislaine Z. Réus
1
, Cecília G. Inácio
1
,
Márcio R. Martins
1
, Clarissa M. Comim
1
and João Quevedo
*,1
1
Laboratório de Neurociências and Instituto Nacional de Ciência e Tecnologia Translacional em Medicina, Programa
de Pós-Graduação em Ciências da Saúde, Unidade Acadêmica de Ciências da Saúde, Universidade do Extremo Sul
Catarinense, 88806-000 Criciúma, SC, Brazil
2
Neurogenetics Section, Neuroscience Department, Centre for Addiction and Mental Health, M5T158 Toronto, ON,
Canada
Abstract: In this study age-, circadian rhythm- and methylphenidate administration- effect on open field habituation and
object recognition were analyzed. Young and adult male Wistar rats were treated with saline or methylphenidate 2.0mg/kg
for 28 days. Experiments were performed during the light and the dark cycle. Locomotor activity was significantly altered
by circadian cycle and methylphenidate treatment during the training session and by drug treatment during the testing
session. Exploratory activity was significantly modulated by age during the training session and by age and drug treatment
during the testing session. Object recognition memory was altered by cycle at the training session; by age 1.5 h later and
by cycle and age 24 h after the training session. These results show that methylphenidate treatment was the major
modulator factor on open-field test while cycle and age had an important effect on object recognition experiment.
Keywords: Memory, methylphenidate, locomotor activity, aging, rat behavior, circadian rhythm.
INTRODUCTION
Attention-deficit/hyperactivity disorder is the most com-
monly diagnosed neuropsychiatry disorder in childhood,
characterized by excessive levels of inattentiveness, impulsi-
vity, and hyperactivity. Most of attention-deficit/hyperacti-
vity disorder (ADHD) patients benefit from treatment with
methylphenidate hydrochloride (MPH), irrespective of the
etiology of the disorder, effectively reducing symptoms in up
to 70% of children patients (Wilens, 2008). However, recog-
nition that ADHD persists into adulthood has led to an
increased use of MPD in adult patients (Quinn et al., 2008).
Studies in humans suggest that blockade of the dopamine
transporter (DAT) produces the therapeutic effect of MPD
by increasing concentrations of dopamine in the central
nervous system (Volkow et al., 1998, 2002). Pharmacologic
properties of MPH have been well characterized in several
preclinical studies: MPH has been found to improve spatial,
working memory, response inhibition, set shifting, and other
prefrontal cognitive functions in children and adults with
attention-deficit/hyperactivity disorder, besides the effects on
attention and impulsivity (Peterson et al., 2008). However,
its mechanism of action is not completely understood (Souza
et al., 2008).
Biological rhythms, particularly circadian rhythm, are an
essential part of the temporal order of living systems and
their disruption has been related with some neuropsychiatric
*Address correspondence to this author at the Laboratório de Neurociências,
PPGCS, UNASAU, Universidade do Extremo Sul Catarinense, 88806-000
Criciúma, SC, Brazil; Fax: #55 48 3443 4817; E-mail: quevedo@unesc.net
Received: July 6, 2009, Revised: August 07, 2009, Accepted: August 25, 2009
disorders. Age related changes have been documented in
endocrine, metabolic and behavioral parameters in a variety
of animal species, including humans. It has been shown that
MPH and other stimulants effects are modulated by age and
circadian rhythms (Gaytan et al., 1996). One of the most
fundamental behaviors an organism exhibits is exploration of
its environment. Exploration includes both spatial explora-
tion and investigation of specific stimuli and is the sum of
motor and locomotor activity, emotional drives (motivation,
anxiety, and fear), and cognitive capacity (Barnett, 1958).
Simple measures of total activity demonstrate that infant rats
are generally more active in the dark than in the light phase
of the daily cycle (Norton et al., 1975), have considerable
locomotor capacity (Moorcroft et al., 1971) and are capable
of rearing (Campbell and Mabry, 1973). As age advances, it
may be expected that the animal behavior performance
decline gradually (Peng et al., 1994). Measures interpreted
as anxiety in rodents include certain uses of the open field
(locomotor - crossings - and exploratory - rearings - activity.
This method is frequently used in adults but is rarely used in
young rats (Smith and Morrell 2007).
Object recognition task has been increasingly used in
recent years as a model for investigation of the neurobiolo-
gical mechanisms of learning and memory, especially
recognition memory that confers the ability to discriminate
between novel and familiar entities (Winters et al., 2008).
Originally developed by Ennaceur and Delacour (1988), this
task can study in a relatively “pure” manner the recognition
memory without the potential complications of interpretation
introduced by, for example, extensive training phases of rule
acquisition, or motivational considerations, because no
explicit reinforcement or aversive stimulation is used during
training (Winters et al., 2008). This is particularly advanta-