Entomologia Experimentalis et Applicata 99: 1–12, 2001.
© 2001 Kluwer Academic Publishers. Printed in the Netherlands.
1
Development and evaluation of an enzyme-linked immunosorbent assay to
detect Pieris rapae remains in guts of arthropod predators
M. A. Schmaedick
1,3
, K. S. Ling
2,4
, D. Gonsalves
2
& A. M. Shelton
1
1
Department of Entomology, NYSAES, Cornell University, Geneva, NY 14456, USA;
2
Department of Plant Pathol-
ogy, NYSAES, Cornell University, Geneva, NY 14456, USA;
3
Current address: Land Grant Program, American
Samoa Community College, PO Box 5319, Pago Pago, AS 96799, USA;
4
Current address: AgriVitis, Inc., 2191 San
Juan-Hollister Road, San Juan Bautista, CA 95045, USA
Accepted: August 31, 2000
Key words: ELISA, polyclonal antisera, sensitivity, specificity, Pieris rapae, predation
Abstract
An enzyme-linked immunosorbent assay (ELISA) was developed to detect remains of Pieris rapae L. (Lepidoptera:
Pieridae) immature stages in the guts of field collected arthropod predators. The assay can be used to help ascertain
the relative importance of arthropod predator species in suppressing P. rapae in cabbage, Brassica oleracea var.
capitata L. The ELISA is sensitive to all immature stages of P. rapae, although first and fifth instars can be
detected more readily than eggs or pupae and third instars showed intermediate detectability. Assays on whole
body homogenates of predators readily detected predation on P. rapae first instars by all seven of the predator
species tested, although response generally declined with increasing predator size. Together the results show that
the P. rapae ELISA possesses a sufficiently high level of sensitivity and specificity to be a useful tool in helping to
elucidate the roles of arthropod predator species in reducing populations of P. rapae in cabbage.
Introduction
Immunological assays to detect prey remains in the
guts of predators collected from the field are a use-
ful tool to help researchers identify important trophic
connections within an ecosystem (Powell et al., 1996).
Once developed, these assays can serve as an efficient
and sensitive means to test large numbers of field col-
lected predators for evidence of feeding on a specific
prey item, such as a key agricultural pest. However,
immunoassays must be carefully assessed before they
can be used in ecological studies (Boreham & Ohiagu,
1978; Sopp et al., 1992; Kidd & Jervis, 1996; Sun-
derland, 1996). Critical aspects that must be evaluated
include sensitivity and specificity, and effectiveness in
detecting predation by different species. This paper
documents these attributes for an immunoassay that
we developed for a key pest of cabbages (Brassica
oleracea variety capitata L.) in New York State.
The target of our immunoassay, the imported cab-
bageworm, Pieris rapae L. (Lepidoptera: Pieridae),
is a serious pest of cabbages and other crucifer crops
in North America and many other parts of the world.
While research elsewhere (including some using im-
munological assays) has shown that predatory arthro-
pods can cause high levels of mortality in P. rapae
populations (Dempster, 1967; Parker, 1970; Ashby,
1974; Hasui, 1977; Jones et al., 1987), until recently
the impact of predators has not been evaluated in New
York. The objective of this study was thus to develop
and characterize an enzyme-linked immunosorbent as-
say (ELISA) to help identify important predators of
P. rapae on cabbages in New York State. In particular,
five aspects of the immunoassay were addressed: (1)
the assay’s sensitivity to first instar P. rapae, the stage
which appears most susceptible to predation (Demp-
ster, 1967; Ashby, 1974; Schmaedick & Shelton,
1999); (2) differences in sensitivity to the different
immature stages of P. rapae; (3) specificity (i.e., the
levels of cross-reactions to other species); (4) ability
to detect predation by seven predator species; and (5)