Oxidative Degradation of Azo Dyes by Manganese Peroxidase under
Optimized Conditions
I. Mielgo, C. Lo ´ pez, M. T. Moreira, G. Feijoo, and J. M. Lema*
Department of Chemical Engineering, Institute of Technology, University of Santiago de Compostela,
E-15782 Santiago de Compostela, Spain
The application of enzyme-based systems in waste treatment is unusual, given that
many drawbacks are derived from their use, including low efficiency, high costs and
easy deactivation of the enzyme. The goal of this study is the development of a
degradation system based on the use of the ligninolytic enzyme manganese peroxidase
(MnP) for the degradation of azo dyes. The experimental work also includes the
optimization of the process, with the objective of determining the influence of specific
physicochemical factors, such as organic acids, H
2
O
2
addition, Mn
2+
concentration,
pH, temperature, enzyme activity and dye concentration. A nearly total decolorization
was possible at very low reaction times (10 min) and at high dye concentration (up to
1500 mg L
-1
). A specific oxidation capacity as high as 10 mg dye degraded per unit of
MnP consumed was attained for a decolorization higher than 90%. Among all, the
main factor affecting process efficiency was the strategy of H
2
O
2
addition. The
continuous addition at a controlled flow permitted the progressive participation of
H
2
O
2
in the catalytic cycle through a suitable regeneration of the oxidized form of the
enzyme, which enhanced both the extent and the rate of decolorization. It was also
found that, in this particular case, the presence of a chelating organic acid (e.g.,
malonic) was not required for an effective operation. Probably, Mn
3+
was chelated by
the dye itself. The simplicity and high efficiency of the process open an interesting
possibility of using of MnP for solving other environmental problems.
Introduction
In recent years the massive increase in the synthesis
of chemicals has led to the production of a wide variety
of compounds, some of which are xenobiotic or persistent
in natural ecosystems. Synthetic dyes belong to this type
of compounds; most of them have an aromatic structure,
which makes them highly stable and thus difficult to
degrade (1). Nowadays, systems based on physicochem-
ical treatments (flocculation, ozonation or membrane
filtration) have currently been used for the treatment of
effluents containing dyes (2). However, these treatments
produce an accumulation of sludges, which will eventu-
ally have to be treated. Conventional biological waste-
water treatment systems, such as activated sludges, are
ineffective in dye removal because most of these com-
pounds pass through these plants without any structural
modification. Legislation relating to colored effluent
discharges is becoming stricter, and new technologies are
needed to solve this environmental problem.
White-rot fungi may constitute an upcoming alterna-
tive for the degradation of dyes and other recalcitrant
compounds because they produce oxidases and per-
oxidases, which are highly oxidative and substrate-
nonspecific. The most outstanding enzymes are lignin
peroxidase (LiP) and manganese peroxidase (MnP), first
described in Phanerochaete chrysosporium (3, 4). The
oxidation of dyes by fungal cultures has been made
possible (5, 6); however, the major limitation associated
with their application relates to the time necessary to
carry out the degradation (7).
From a practical point of view, the use of peroxidases
in vitro for that purpose may represent a more feasible
system, provided that a number of conditions are ful-
filled: (a) a simple and applicable system; (b) minimal
requirements of compounds; (c) low cost and stable
enzyme; and (d) short treatment periods. This system
could be implemented as a pre- or as a post-treatment
for conditioning those concentrated fractions of effluents
with strong color.
There are several studies of in vitro incubations of azo
dyes with crude or purified LiP (8-11) or MnP (12). The
assays reported were performed on a very small scale,
and only a limited decolorization yield was achieved. This
low efficiency may be due to either some compounds
lacking in the amounts required or to the nonoptimized
physicochemical conditions, both of which are required
for the enzymatic action.
The main objective of this paper is the evaluation of
the use of MnP from Bjerkandera sp. BOS55 for the
degradation of azo dyes. This enzyme presents high
versatility since it has both MnP and LiP substrate
interaction sites (7, 13), and in this way, it arises as an
interesting alternative to develop an in vitro degradation
system. The action of MnP depends on the combined
action of several compounds, referred to as substrates,
cofactors and mediators, which initiate, participate in,
and allow the completion of the catalytic cycle. Therefore,
special attention was paid to study the influence of the
* To whom correspondence should be addressed. Tel: 34-981-
563100, Ext. 14231. Fax: 34-981-595012. E-mail: jmlema@usc.es.
325 Biotechnol. Prog. 2003, 19, 325-331
10.1021/bp020136w CCC: $25.00 © 2003 American Chemical Society and American Institute of Chemical Engineers
Published on Web 01/10/2003