© 2002 Blackwell Science Ltd
Microenvironments and microbial community structure
in sediments
water interface. Microbial mats are laterally compressed
ecosystems in surface sediments, which support most of
the major biogeochemical cycles within a vertical dimen-
sion of only a few millimetres (Paerl and Pinckney, 1996).
Consequently, the understanding of interactive influences
between oxygen, sulphide, manganese, iron, redox po-
tential, pH and microorganisms requires microtechniques
capable of probing these interactions. We describe the
application of such techniques to the sediment–water
interface of Sælen lake, a semi-enclosed basin connected
to the fjord system of the Norwegian coast.
Sælen lake is a meromictic lake and exhibits a wide
variety of oxygen conditions, from seasonal fluctuations
in the shallow parts to permanent anoxia in the central
basin. One notable advantage of using such an environ-
ment, is that the burrowing activities of macro-organisms
is minimal because few of these organisms survive anoxic
conditions and, consequently, only chemical and micro-
biological reactions can contribute to the chemical fluxes
between sediment and the water column. The microbial
ecology of this ‘model environment’ has been extensively
studied previously (Indrebø et al., 1979a; b; Heldal et al.,
1996; Øvreås et al., 1997). The aim of this study was to
explore the potential of a combined chemical and micro-
biological approach as part of a study of organic carbon
oxidation processes in Saelen lake sediments. We were
particularly interested in examining the participation of the
microbial mat community in the Fe and Mn cycling at the
sediment–water interface.
Results
Chemical characterization of the biofilm in
surface sediment
In October 2000, O
2
, H
2
S, pH and redox potential were
measured across the sediment–water interface of Sælen
lake and data corresponded to daylight conditions. The
depth profiles are shown in Fig. 1A and B. Oxygen satu-
ration increased from 45% in bottom water to 50% just
above the sediment–water interface (Fig. 1A) and then
decreased to 0% saturation at 1 mm depth into the sedi-
ment (Fig. 1B). There was a 0.2 mm overlap between
the O
2
and H
2
S profiles. pH showed a maximum of 7.53
near the sediment–water interface followed by a steep
decrease just below. The redox potential was relatively
high (+500 mV) in bottom water and in the 0–0.9 mm zone
Environmental Microbiology (2002) 4(2), 97–105
S. P. C. Tankéré,
1
* D. G. Bourne,
1
F. L. L. Muller
2
and
V. Torsvik
1
Departments of
1
Microbiology and
2
Chemistry,
University of Bergen, Bergen, Norway.
Summary
The aim of this study was to explore the potential of
a combined chemical and microbiological approach
as part of a study of organic carbon oxidation pro-
cesses in sediments. An assessment of microbiologi-
cal diversity using molecular techniques was carried
out in combination with high resolution chemical
measurements at the sediment–water interface of a
coastal lagoon affected by eutrophication in autumn
2000. There was a 0.2 mm overlap between the O
2
and
H
2
S profiles. pH showed a maximum just above the
sediment–water interface coinciding with an oxygen
maximum, suggesting photosynthetic activity, and a
minimum coinciding with the O
2
–H
2
S interface. The
redox potential was high in bottom water and sur-
face sediment, reflecting the presence of oxygen and
oxides, and reached low values after a step-wise
decrease at –18 mm. Reduction of Fe occurred within
the biofilm at the O
2
–H
2
S interface and was mostly
due to reduction by H
2
S. The elevated concentrations
of dissolved Mn in the oxic water may have been
caused either by in situ production within organic
aggregates or lateral water flow from sites nearby at
which Mn
2+
diffuses out of the sediment. Sequences
related to sulphur chemolitotrophs were retrieved
from the biofilm samples, which is consistent with the
small overlap between O
2
and H
2
S observed in this
biofilm. Although the resolution of techniques used
was different, sequencing results were consistent
with chemical data in delineating the same horizons
according to redox, pH or ecological properties.
Introduction
Aquatic systems that receive large inputs of organic mate-
rial, as frequently occurs in lakes, rivers, estuaries and
coastal waters affected by eutrophication, are charac-
terized by marked chemical changes in the sediment–
*For correspondence. E-mail sophie.muller@im.uib.no (S.P.C.
Tankéré); Tel. (+47) 55584638 Fax (+47) 55589671.