African Journal of Plant Science Vol. 3 (4), pp. 064-073, April 2009
Available online at http://www.academicjournals.org/AJPS
ISSN 1996-0824 © 2009 Academic Journals
Full Length Research Paper
Investigation of morphophysiological variation in field
bindweed (Convolvulus arvensis L.) populations of
Karaj, Varamin, and Damavand in Iran
A. Mehrafarin
1
, F. Meighani
2
, M. A. Baghestani
2
, M. J. Mirhadi
1
and M. R. Labbafi
2
*
1
Department of Weed Science, Faculty of Agriculture, Science and Research Campus, Islamic Azad University, Tehran,
Iran.
2
Weed Research Department, Iranian Plant Protection Research Institute, P. O. Box, 19395-1454, Tehran, Iran.
Accepted 19 March, 2009
Diversity in field bindweed (Convolvulus arvensis L.) populations collected from Karaj, Varamin, and
Damavad during 2006 (for seed collection) to 2007 (for seed germination and plant growth in green-
house) at the Weed Research Department, Iranian Plant Protection Research Institute for identification
of morphophysiological variation using multivariate analysis methods. The most important variables
were shoot dry weight, leaf dry weight, and leaf area, respectively. Results showed 11, 15 and 16
biotypes in Karaj, Varamin, and Damavand populations, respectively. Varamin was clustered near to
Damavand, but both these populations had significant differences with Karaj population.
Key words: Biodiversity, ecotype, biotype, principal component analysis, cluster analysis, field bindweed,
Convolvulus arvensis.
INTRODUCTION
Field bindweed (Convolvulus arvensis) is one of the 10
noxious weeds in the world and 54 countries reported
that it is found as a weed in 32 different crops (Swan,
1980; Holm et al., 1991). This is considered as one of the
problematic weeds in Iran (Shimi and Termeh, 2004) and
found in a wide range of habitats (Weaver and Riley,
1982). This species have a broad geographical range,
often include either ecotypes or biotypes (Klingaman and
Oliver, 1996). It is self-incompatible species which may
play an important role in maintaining the high degree of
phenotypic variation (Westood, and Weller, 1997).
Numerous examples of intraspecific variation in growth
and morphology of weed species have been reported
(Degennaro and Weller, 1984). Morphological parame-
ters are generally used as a tool for investigation of diver-
sity and genetic relatedness (Hubner et al., 1998). Seve-
ral morphologically distinct biotypes of field bindweed
have been identified and more than one biotype often
exists in the same infested area. The morphological
variability in the species is thought responsible for diff-
erential response of field bindweed to herbicides (Duncan
and Weller, 1987). Variation in morphology and herbicide
*Corresponding author. E-mail: mohammad1700@yahoo.com.
susceptibility of field bindweed has been observed by
various researchers (Degennaro and Weller, 1984).
The objective of this investigation is to determine the
differences in morphophysiological characteristics of field
bindweed biotypes or ecotypes in these regions.
MATERIALS AND METHODS
Seed of field bindweed (C. arvensis L.) were collected from Karaj,
Varamin, and Damavand in Tehran province in September 2006. In
March 2007, after seed scarification with sulfuric acid 97%, these
were planted in plastic pots having mixtures of sterilized clay, sand,
manure and perlite in a ratio of 1:5:5:0.5, respectively. The plants
were maintained in a greenhouse for 22 wks under day/night temp-
eratures of 30/18 ± 4ºC and 500 μmol.m
-2
.s
-1
with supplemental
lighting (incandescent and fluorescent) to provide a 14 h day length
with 45% relative humidity. Plants were irrigated weekly, and as
needed with a nutrient solution containing 200 ppm N, 100 ppm P,
and 100 ppm K (Samadani and Minbashi, 2004).
Morphophysiological characteristics studied
At the end of flowering (154 days after sowing), 60 plants were
randomly collected from each population and observations were
recorded on traits like. shoot number, shoot fresh weight, shoot dry
weight, shoot water content, stem dry weight, leaf dry weight, root
dry weight, whole plant dry weight, whole plant biomass, collar
(crown) diameter, leaf number, leaf area, chlorophyll concentration,