Psychopharmacology (1994) 115:454462
Psychopharmacolog
© Springer-Vertag 1994
Reversal of stress-induced anhedonia by the dopamine recep
agonist, pramipexole
Paul Willner 1, Spiros Lappas, Survjit Cheeta ~, Richard Muscat 2
Department of Psychology, City of London Polytechnic, Old Castle St, London E1 7NT, UK
Received: 19 July 1993 / Final version: 16 December 1993
Abstract. Chronic exposure to mild unpredictable stress
has previously been found to depress the consumption of
a palatable (1%) sucrose solution, and to attenuate food°
induced place preference conditioning. In this study the
effects of pramipexole (SND-919), a dopamine D2 ago-
nist, were studied during 7-9 weeks of chronic treatment.
Pramipexole (1.0 mg/kg per day) reversed the suppres-
sion of sucrose intake in stressed animals, increasing su-
crose intakes above the levels seen in untreated non-
stressed controls. Pramipexole also increased sucrose in-
take in nonstressed animals; these effects were accompa-
nied by increases in water intake and tended to correlate
with weight loss. Drug-treated stressed animals also lost
weight, but in this case water intake was unaffected. A
second group of animalsreceived a higher dose of
pramipexole (2.0 mg/kg per day). The effects of the two
doses were very similar. After three weeks of treatment,
these animals were switched to a lower dose of pramipex-
ole (0.1 mg/kg per day). Increases in sucrose intake were
maintained over three weeks of treatment at the lower
dose, with significant recovery of body weight. Two fur-
ther groups received the same doses of pramipexole
(1.0 mg/kg for 6 weeks or 2.0 mg/kg for 3 weeks followed
by 0.1 mg/kg thereafter), but received intermittent (twice-
weekly) drug treatment. Intermittent pramipexole treat-
ments also tended to increase sucrose intakes, but the
results were less consistent from week to week. Following
6-8 weeks of pramipexole treatment, food-induced place
preference conditioning was studied in all animals. Un-
treated stressed animals showed no evidence of place
conditioning. Normal conditioning was seen in both
groups of stressed animals treated daily with pramipex-
ole (at 1.0 and 0.1 mg/kg) and in the group treated twice
weekly at the higher dose (1.0 mg/kg); intermittent treat-
ment at the lower dose (0.1 mg/kg) was ineffective. The
1present address: Dept of Psychology, University of Wales,
Swansea, Singleton Park, Swansea SA2 8PP, UK
2 Present address: Dept of Biomedical Sciences, University of Malta,
Msida, Malta
Correspondence to: P Willner
results indicate that pramipexole exerts rapid anti-an-
hedonic effects in the chronic mild stress model. This
conclusion is complicated, but not undermined, by drug~
induced weight loss and by the presence of significa
drug effects in nonstressed control animals.
Key words: Stress Sucrose drinkingPlace preference
conditioning -Reward -Pramipexole - D2 agonist -
Rats
Chronic sequential exposure to mild unpredictable stress
(chronic mild stress: CMS) causes, in rats, a generalize
decrease in responsiveness to rewards,which may
provide an animal model of anhedonia, the pervasive loss
of pleasure capacity characteristic of melancholic depres
sions (Klein 1974; Fawcett et al. 1983). Effects of CMS
include decreased consumption of, and preference for
palatable sweet solutions (Willner et al. 1987; Muscat
and Willner 1992), attenuation of the rewarding proper-
ties of drugs (amphetamine, morphine) and natural rein-
forcers (sweet solutions, food) in the place preference
paradigm (Papp et al. 1991, t992), and increases in th
threshold for intracranial self-stimulation (Moreau et al.
t992, 1994). All of these effects may be reversed by chron
ic treatment with antidepressants, including tricyclics
(Willner et al. 1987; Muscat et al. 1990; Sampson et al
1991; Moreau et al. 1992), monoamine oxidase inhibitors
(Moreau et al. 1994), and atypical antidepressants, suc
as specific noradrenatine or serotonin reuptake inhibitors
(Muscat et al. 1992a) and mianserin (Cheeta et al. 1992
CMS causes abnormalities of dopamine (DA) neuro-
transmission within the nucleus accumbens (Willner et al
1991a; Stamford et al. 1991) and a decreased sensitiv
to behavioural effects of DA agonists (Papp et al. 1991
1993a). A functional role for DA in the behavioural ef
fects of CMS is indicated by the fact that antidepressant
effects in the CMS model are reversed by acute adminis-
tration of DA receptor antagonists (Muscat et al. 1990
1992a; Sampson et al. 1991), which suggests that th