4/13/2016 C:\DOCS\MSS\PIECH.HTM
http://cogprints.org/68/1/piech.htm 1/7
Pergamon
Physiology & Behavior. Vol. 57, No. 2, pp. 209212, 1995
Copyright © 1995 Elsevier Science Ltd
00319384(94)002754
Ingestion of Amniotic Fluid by Postpartum Rats
Enhances Morphine Antinociception Without
Liability to Maternal Behavior
J. A. TARAPACKI, M. PIECH AND M. B. KRISTAL
1
Behavioral Neuroscience Program, Department of Psychology, State University of New York at Buffalo,
Buffalo, NY 142604110 email: KRISTAL @ acsu.buffalo.edu
Received 1 April 1993
TARAPACKI, J. A., M. PIECH AND M.B. KRISTAL. Ingestion of amniotic fluid by postpartum rats enhances
morphine anti nociception without liability to maternal behavior. PHYSIOL BEHAV 57(2) 209212. 1995.
Ingestion of amniotic fluid or placenta by rats has been shown to enhance opioidmediated analgesia induced by
morphine injection, foot shock, vaginal/cervical stimulation, or late pregnancy. The present study was designed to
determine whether this mechanism might be a means of providing greater analgesia during the periparturitional
period without contributing to the disruption of maternal behavior (measured primarily as retrieval) that can result
from excessive opioid levels. Postpartum primiparous rats, injected with either 2 or 3 mg/kg morphine sulfate or
vehicle and given orogastric infusions of either amniotic fluid or saline, were tested for maternal behavior. Pain
threshold (determined by tailflick latency test) in rats injected with 2 mg/kg morphine and infused with amniotic
fluid was elevated to a level that did not differ significantly from that of a separate group of rats injected with 3
mg/kg morphine and infused with saline. This enhanced analgesia was not, however, accompanied by the
significant disruption of maternal behavior found among the rats receiving the higher morphine dose.
Parturition
Maternal behavior
Opioids
Analgesia
Morphine
Placentophagia
Pain
POEF
Tailflick test Amniotic
fluid
AMONG most nonaquatic placental mammals, placentophagia, the ingestion of birth membranes and fluids, is typical
during delivery (16,17). One of the effects of this behavior is the enhancement of opioidmediated analgesia (16). By
itself, the ingestion of placenta or amniotic fluid does not produce analgesia (20,23), nor does it appear to enhance
nonopioidmediated analgesia (19). It does, however, enhance analgesia produced by injection of morphine (1,6,18
20,22,23,31), foot shock (22), vaginal/cervical stimulation (1,7,23), and late pregnancy (21). The relevance of this
phenomenon to normal parturition is supported by findings that opioid levels (8,9,12,14,26,27,34) and pain thresholds
(2,10,11) are elevated during pregnancy and show peaks at or near delivery.
One question that this strategy of potentiating the analgesic effect of the opioids available at parturition raises is whether
it offers some advantage over increased opioid production or release. The possibility has been suggested (16) that it may
be a way of providing the benefit of greater analgesia with less risk of the disruption of maternal behavior that a number
of studies (4,13,15,24,25,29) have shown to be a consequence of excessive levels of opioids. Those studies used
retrieval and grouping of pups as a combined index of the quality of maternal behavior. The choice of these criteria is
supported by the observation that the test rats exhibiting these components of maternal behavior, especially retrieval,
also exhibit most if not all the other components of maternal behavior (e.g., crouching over pups in a nursing posture;