PATHOLOGY ONCOLOGY RESEARCH Vol 13, No 4, 2007
Article is available online at http:/]www.webio.hu/por/2007/13/4/0336
ARTICLE
Expression of Connective Tissue Growth Factor (CTGF/CCN2)
in a Mouse Model of Rhabdomyosarcomagenesis
Stefania CROCI, ~Lorena LANDUZZI, 2 Giordano NICOLETTI, 2 Arianna PALLADINI, 1 Agnese ANTOGNOLI, l
Carla DE GIOVANNI, 1 Patrizia NANNI, ~Pier-Luigi LOLLINI t
~Cancer Research Section, Department of Experimental Pathology, University of Bologna;
2Laboratorio di Ricerca Oncologica, Istituti Ortopedici Rizzoli, Bologna, Italy
Connective tissue growth factor (CTGF/CCN2) is a
cysteine-rich matricellular protein that belongs to
the CCN (CYR61, CTGF, __NOV)protein family. It is
highly expressed by human rhabdomyosarcoma
cells and sustains their survival. In this study we
investigated CCN2 expression in a mouse model of
spontaneous rhabdomyosarcomagenesis that com-
bines HER-2/neu oncogene activation and p53
oncosuppressor gene inactivation (BALB-p53neu
mice). Murine rhabdomyosarcoma cells showed a 4-
26 fold increase in CCN2 mRNA expression regard-
ing to normal thigh muscle. Moreover, they
expressed CCN2 protein at levels comparable to
human rhabdomyosarcoma cells. Therefore BALB-
p53neu mice might be useful for the evaluation of
the role played by CCN2 in rhabdomyosarcoma in
vivo. (Pathology Oncology Research Vol 13, No 4,
336-339)
Key words: rhabdomyosarcoma, mouse model, CCN proteins, CCN2]CTGF
Introduction
Connective tissue growth factor (CTGF or CCN2)
belongs to the CCN family of proteins that comprises
other five members: CCN1/CYR61 (cysteine-rich protein
61), CCN3]NOV (nephroblastoma overexpressed gene),
CCN4/WISP-1 (Wnt-induced secreted protein-l),
CCN5/WISP-2 and CCN6]WISP-3. z They are secreted,
cysteine-rich, matricellular proteins that share a common
modular structure of four domains: insulin-like growth
factor binding domain, von Willebrand type C domain,
thrombospondin-1 domain and cystine knot domain (with
the exception of CCN5 that lacks the last one). 2 CCN2 has
multiple biological functions, dependent on the cell type
and the cellular context. It regulates cell proliferation, sur-
vival, migration and differentiation, and has a role in
angiogenesis, wound healing, fibrosis, chondrogenesis
and osteogenesis. 8 Moreover, all CCN proteins are emerg-
ing as key regulators of tumorigenesis, involved in the
Received: Dec 4, 2006; accepted: Nov 10, 2007
Correspondence: Prof. Pier-Luigi LOLLINI, PhD, Section of Can-
cer Research, Department of Experimental Pathology, University of
Bologna, Viale Filopanti 22,1-40126, Bologna, Italy. Tel.: +39-051 -
2094786, Fax: +39-051-242169, E-mail: pierluigi.lollini@unibo.it
regulation of tumor cell growth and tumor-stroma interac-
tions, thus being promising targets for cancer therapy. 9
We have previously found that CCN2 might be a therapeu-
tic target for human rhabdomyosarcoma, the most common
soft tissue sarcoma of skeletal muscle origin in childhood.
CCN2 was found highly expressed by human rhabdomyosar-
coma cell lines and tumor specimens of embryonal and alve-
olar histotypes. Moreover, functional studies have indicated
that it sustained human rhabdomyosarcoma cell survival and
myogenic differentiation in an autocrine way.3 In this study
we took advantage of a mouse model of rhabdomyosarcoma-
genesis established in our laboratory through the combination
of HER-2/neu oncogene activation and p53 oncosuppressor
gene inactivation (BALB-p53neu mice).6 We investigated
whether murine rhabdomyosarcoma cells expressed CCN2
like human rhabdomyosarcoma cells making BALB-p53neu
mice a suitable preclinical model for the study of the role
played by CCN2 in rhabdomyosarcoma in vivo.
Materials and methods
Mice and cell lines
BALB/c mice transgenic for the activated rat HER-2/neu
oncogene under the control of the MMTV-LTR promoter
and with a knockout allele for the p53 oncosuppressor gene,
© 2007 Ar~nyi Lajos Foundation