Introduction
Insulin-like growth factor-I (IGF-I) is a naturally
occurring polypeptide produced in the liver, muscle and fat
tissues. IGF-I is present in the circulating blood supply,
usually bound with 1 of 6 binding proteins (IGFBPs). These
binding proteins form stable complexes with IGF-I that act to
maintain IGF-I in circulation, and additionally control the
distribution, function and activities of IGF-I (Hossner et al.
1997). IGF-I influences growth and development during the
postnatal growth period through endocrine, autocrine and
paracrine mechanisms. IGF-I subsequently mediates the
effects of growth hormone (GH), has localised effects on
tissues at or close to sites of production, and also acts as an
insulin-like metabolic regulator (Hossner et al. 1997). IGF-I
and IGF-II, through their interactions with hormones and
binding proteins, form part of a very complex system; many
aspects of which are not completely understood.
The concentration of circulating IGF-I can be measured
relatively easily in blood plasma or serum with appropriate
assay techniques (see Holly and Cwyfan Hughes 1994). As a
result of its relative ‘ease’ of measurement compared with
GH (i.e. serial measurements are not required), IGF-I was
historically used as a substitute marker for GH in breeding
experiments. Blair et al. (1988) demonstrated that divergent
selection for IGF-I resulted in changes to both IGF-I
concentrations and growth performance in rodents.
Moderate estimates of heritabilities for IGF-I were later
reported for cattle (Davis and Bishop 1991; Herd et al. 1995;
Johnston et al. 2001, 2002) and pigs (Lamberson et al. 1995;
Luxford et al. 1998a; Lahti et al. 2001; Cameron et al. 2003;
Suzuki et al. 2004), implying utility for selection purposes in
domestic livestock species. Single trait divergent selection
lines for IGF-I are currently under development for cattle
and sheep (Davis and Simmen 1997; Blair et al. 2002) but
animal numbers are relatively low. While there are several
studies that confirm circulating IGF-I concentrations are
usually positively correlated with growth status in laboratory
and livestock species, this is not always the case. It should
also be noted that the majority of work reported on IGF-I has
not been conducted on very juvenile animals, as is the case
considered for trials here.
In growing animals, Hossner et al. (1997) noted that
increased growth in rats due to GH administration did not
increase circulating levels of IGF-I, although IGF-I
concentrations in hepatic and renal tissues were increased.
Dunaiski et al. (1997) further showed that the infusion of
Australian Journal of Experimental Agriculture, 2005, 45, 783–792
0816-1089/05/080783 10.1071/EA05048 © CSIRO 2005
K. L. Bunter
A,C
, S. Hermesch
A
, B. G. Luxford
B
, H-U. Graser
A
and R. E. Crump
A
A
Animal Genetics and Breeding Unit, University of New England, Armidale, NSW 2351, Australia.
B
QAF Meat Industries, PO Box 78, Redlands Road, Corowa, NSW 2646, Australia.
C
Corresponding author. Email: kbunter2@une.edu.au
Abstract. Insulin-like growth factor-I (IGF-I) is a naturally occurring polypeptide produced in the liver, muscle
and fat tissues. It is known to be associated with growth and development during the postnatal growth period.
Evidence for strong genetic correlations between juvenile IGF-I and performance traits would suggest this
physiological measure would be useful as an early selection criterion. This paper reports estimates of genetic
parameters from 9 trials where IGF-I was measured in juvenile pigs. All trials involved populations undergoing
active selection for improved performance (e.g. efficient lean meat growth). Juvenile IGF-I was moderately
heritable (average h
2
: 0.31) and influenced by common litter effects (average c
2
: 0.15). Genetic correlations (r
g
)
between juvenile IGF-I and backfat (BF), feed intake (FI) or feed conversion ratio (FCR) traits were generally large
and positive: r
g
averaged 0.57, 0.41 and 0.65, respectively. Phenotypic correlations (r
p
) between juvenile IGF-I and
BF, FI or FCR were much lower (r
p
averaged 0.21, 0.09, and 0.15, respectively) as residual correlations between
IGF-I and these performance traits were low, consistent with being measured at very different times. Correlations
(genetic or phenotypic) between juvenile IGF-I and growth traits (e.g. lifetime daily gain or test daily gain) were
relatively low, with average values within ± 0.09 of zero. Results from the trials reported here, and several
physiological studies, indicate that information on juvenile IGF-I concentration can be used as an early
physiological indicator of performance traits traditionally measured later in life. There is a clear role for juvenile
IGF-I to facilitate pre-selection and more accurate selection of livestock for hard to measure traits, such as FCR, in
pig breeding programs.
Insulin-like growth factor-I measured in juvenile pigs is genetically
correlated with economically important performance traits
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