© 2008 The Authors. Journal compilation © 2008 OEPP/EPPO, Bulletin OEPP/EPPO Bulletin 38, 507–509 507
Blackwell Publishing Ltd
First record of Phytomyza gymnostoma Loew
(Diptera: Agromyzidae) a leaf mining pest of leeks in Greece
K. B. Simoglou
1
, E. Roditakis
2
, M. Martinez
3
and N. E. Roditakis
2
1
Prefecture Agriculture Directorate of Heraklion, Department of Plant Protection, Certification & Quality Control, Evans 60, 71201 Heraklion,
Greece
2
Plant Protection Institute of Heraklion, National Agricultural Research Foundation, 71003 Heraklion, Greece; e-mail: e-roditakis@her.forthnet.gr
3
Institut National de la Recherche Agronomique, UMR CBGP, 2 place Pierre Viala (Bâtiment 16), 34060 Montpellier, Cedex 01, France
In autumn 2007, an allium leafminer, Phytomyza gymnostoma Loew (Diptera: Agromyzidae), was
recorded for the first time in non-commercial leek crops in the areas of Elassona and Katerini in
Central Greece. Leek plants were severely damaged by the leafminer larvae infestation. Typical
feeding symptoms were observed, caused by the mining behaviour of larvae within inner and
outer leaves of leek plants, resulting in the formation of descending galleries. Pest control was
performed locally using commercial insecticides with unknown efficacy mainly due to ignorance
of the species biology.
Introduction
Onion (Allium cepa L.), leek (Allium porrum L.) and garlic
(Allium sativum L.) are the most important allium crops in
Greece. In 2006, a total of 13 000 ha were cultivated mainly in
Northern and Central Greece (mostly in areas of Thessaloniki,
Larissa, Pella and Trikala) (onions: 9200, leeks: 2100, garlic:
1700 ha). Approximately 213 thousand tonnes of onion (green
and dry), 44 thousand tonnes of leeks and 13 thousand tonnes
of garlic were produced. In order to cover the local consumption
Greece also imports allium crop products. Imports in 2005 were
estimated around 8 million USD. Greek exports are limited
(garlic and onions, 680 000 USD), mainly to European countries
and US (FAO, http://faostat.fao.org/).
Allium crops have many pests, the most important being the
onion thrips (Thrips tabaci Lindeman, Thysanoptera: Thripidae),
the onion maggot (Delia antiqua (Meigen), Diptera: Anthomyiidae)
the leek or onion moth (Acrolepiopsis assectella (Zeller),
Lepidoptera: Acrolepiidae) and the leafminers (Diptera:
Agromyzidae) (Alford, 2000). There are 7 leafminer species
that have been reported as pests of allium crops. However there
are only three species reported as pests of leek; Liriomyza
chinensis (Kato), Liriomyza nietzkei Spencer and Phytomyza
gymnostoma Loew. All three species have been reported in
Europe (Dempewolf, 2004).
The Allium leafminer P. gymnostoma (formerly Napomyza
gymnostoma) is a pest of leek (A. porrum) and chives (A.
schoenoprasum L.) and to a lesser extent of onion (A. cepa),
garlic (A. sativum), shallot (A. ascalonicum L.) and of ornamental
Allium plants (Agallou et al., 2004). The pest is causing
extensive damage to crops in an increasing number of European
countries. It has been reported as an Allium pest from Central,
Eastern and more recently Western Europe, as well as in Asia
from Turkey and Turkmenistan (Seljak, 1998; Zandigiacomo
et al., 2002; Bouchery & Martinez, 2004; Collins & Lole, 2005;
EPPO Reporting Service, 2005; MacLeod, 2007; Smith et al.,
2007).
The Allium leafminer larvae mine the leaves and bulbs,
making the plants soft and susceptible to bacterial and fungal
infections, such as white rot (Sclerotium cepivorum Berk).
Heavy infestations completely destroy plants (Agallou et al.,
2004; Collins & Lole, 2005).
Adult flies are about 3 mm long, with a largely yellow head,
black and yellow areas underside and dark legs with yellow
joints between the femur and tibia. It has two generations per
year in Central Europe. Adults emerge in the spring from
pupae that overwintered within host plants. Females lay eggs at
the base of host plant stems. The larvae of the first (spring)
generation develop in garlic and onion stems and bulbs. First
generation pupae diapause during the summer before adults
emerge and lay eggs in leeks or other Allium species in the
autumn, for the second generation. Pupae of the second
generation overwinter in host plants, before adults emerge the
following spring to lay eggs on hosts such as onions and garlic
(MacLeod, 2007; EPPO Reporting Service, 2005).
P. gymnostoma was added to the EPPO Alert List by the
EPPO Secretariat in April 2005 following reports in the scientific
literature that the organism was spreading within the EPPO
region and becoming a significant pest of Allium crops (EPPO
Reporting Service, 2005). However, at the 38th meeting of the
EPPO Panel on Phytosanitary Measures, P. gymnostoma was
removed from the Alert List
1
(EPPO Secretariat, 2006).
1
Editor’s note: the reason P. gymnostoma was removed was that it was
widespread in the EPPO region.