Fish as Bioindicators to Assess the Effects of Pollution in
Two Southern Brazilian Rivers Using the Comet Assay
and Micronucleus Test
Vanessa Moraes de Andrade,
1
* Juliana da Silva,
1,2
Fernanda R. da Silva,
1
Vanina D. Heuser,
1
Johnny F. Dias,
3
Maria Lu ´cia Yoneama,
4
and Thales R.O. de Freitas
1
1
Departamento de Gene ´tica, Instituto de Biocie ˆncias, Universidade Federal do Rio
Grande do Sul, Porto Alegre, Rio Grande do Sul, Brazil
2
Laborato ´rio de Gene ´tica Toxicolo ´gica, PPG em Ensino de Cie ˆncias e Matema ´ tica,
Universidade Luterana do Brasil, Canoas, Rio Grande do Sul, Brazil
3
Laborato ´rio de Implantac ¸a ˜o Io ˆnica, Instituto de Fı ´sica da Universidade Federal do
Rio Grande do Sul, Porto Alegre, Rio Grande do Sul, Brazil
4
Programa de Po ´ s-Graduac ¸a ˜o em Geologia, Universidade do Vale do Rio dos
Sinos, Sa ˜o Leopoldo, Rio Grande do Sul, Brazil
Industrial effluents, agricultural runoff, and municipal
wastewaters contain unknown substances and com-
plex mixtures that are released into the environment
and can lead to contamination of surface and subsur-
face waters. In the present report, we have used the
alkaline Comet assay and the micronucleus (MN) test
to detect the genotoxicity due to multiple sources of
pollution in the peripheral blood of two native estua-
rine fish (mullet and sea catfish) and evaluated possi-
ble interactive genotoxic effects from multiple contam-
inants and the seasonal variation of the genotoxicity.
Mullet and sea catfish were captured in the Traman-
daı ´ and Mampituba Rivers in the southern Brazilian
state of Rio Grande do Sul. Reference animals were
obtained from the Armaze ´m lagoon. Fish captured in
the two estuaries during the four seasons over a pe-
riod of 2 years had increased levels of DNA damage
and MN frequencies relative to the reference fish. In
general, the alkaline Comet assay was more sensitive
to the genotoxicity of the river contaminants than the
MN test. The Comet assay demonstrated significant
differences in fish captured at different seasons and at
the two river sites, while the MN test showed signifi-
cant differences only for the annual average for mullet
from both sites and fish from the control site. The
increases in DNA damage appear to be related to the
increase in the number of people in the towns close to
the study areas during the warm spring and summer
seasons. Although no specific cause-effect relation-
ships were established, comparison of the chemical
contaminants and physical variations in the rivers with
the genotoxicity data indicate that there may be some
association between hydrocarbons, metals, pH, and
water temperature and the level of damaged cells
observed in mullet and sea catfish from the Tramandaı ´
and Mampituba estuaries. Environ. Mol. Mutagen.
44:459 – 468, 2004. © 2004 Wiley-Liss, Inc.
Key words: fish; Comet assay; micronucleus test; mullet (Mugil sp.); sea catfish (Netuma sp.);
biomonitoring
INTRODUCTION
Industrial effluents, agricultural runoff, and municipal
wastewaters contain unknown substances and complex mix-
tures that are released into the environment and can lead to
contamination of surface and subsurface waters [Houk,
1992; Kale et al., 1995; White et al., 1996]. Compounds
present in polluted water are capable of causing biological
alterations that can affect particular populations and entire
ecosystems [Rajaguru et al., 2003]. Organisms inhabiting
areas influenced by effluent discharges can suffer genetic
damage, and humans using polluted water are at risk of
similar genotoxic effects and cancer development [Stahl,
1991; De Flora et al., 1993]. The continued production and
release of pollutants into the aquatic environment have
made investigating the genotoxic potential of inland and
coastal waters a major task of environmental pollution-
control monitoring [Rajaguru et al., 2002]. Although it has
Grant sponsor: Coordenac ¸a ˜o de Aperfeic ¸oamento de Pessoal de Nı ´vel
Superior (CAPES).
*Correspondence to: Vanessa Moraes de Andrade, Departamento de Ge-
ne ´tica, Instituto de Biocie ˆncias, Universidade Federal do Rio Grande do
Sul, Av. Bento Gonc ¸alves 9500, CP 15053, Porto Alegre, RS, 91501-970,
Brazil. E-mail: vmoraesdeandrade@yahoo.com.br
Received 20 April 2004; provisionally accepted 5 June 2004; and in final
form 13 August 2004
DOI 10.1002/em.20070
Published online 29 October 2004 in Wiley InterScience (www.
interscience.wiley.com).
Environmental and Molecular Mutagenesis 44:459 – 468 (2004)
© 2004 Wiley-Liss, Inc.