[CANCER RESEARCH 43,1945-1950, May 1983]
0008-5472/83/0043-OOOOS02.00
Chemical Enhancement of Viral Transformation in Syrian Hamster Embryo
Cells by Gaseous and Volatile Chlorinated Methanes and Ethanes1
George G. Hatch,2 Patricia D. Mamay, Mark L. Ayer, Bruce C. Casto, and Stephen Nesnow
Northrop Services, Inc., Research Triangle Park, North Carolina 27709 [G. G. H., P. D. M., M. L A., B. C. C.], and Carcinogenesis and Metabolism Branch, Genetic
Toxicology Division, United States Environmental Protection Agency, Research Triangle Park, North Carolina 27711 [S. N.]
ABSTRACT
Methods were developed for exposing cells in vitro to gases
or vapors of volatilized organic liquids. Compounds were se
lected for their industrial importance, environmental impact, and
suspected role in the etiology of some human cancers. Exposure
chambers were designed for easy insertion of dishes of cultured
cells and were equipped with inlet and outlet ports for introduc
tion and purging of test gases. A gas delivery system utilizing a
mass flow meter was used for the quantitative distribution of
test gases into exposure chambers. For volatile compounds,
appropriate volumes of cold (4°)liquids in glass Petri dishes
were quickly placed into chambers, the system sealed, and the
compounds rapidly volatilized at 37°. For exposure, the cells
and chambers were placed in an incubator and rocked at a
constant rate so that a portion of the cells was always in direct
contact with the test gases or vapors. Known sample volumes
were removed after various treatment times and test gas con
centrations determined by standard gas Chromatographie tech
niques. After exposure, the cells were removed and assayed for
viability and increased sensitivity to viral transformation. Under
these experimental conditions, the volatile liquids 1,1,1-trichlo-
roethane, dichloromethane, chloroform, 1,2-dichloroethane, and
1,1-dichloroethane significantly enhanced transformation of Syr
ian hamster embryo cells by SA7 adenovirus, while acetone
exerted no effect. The gases chloromethane and vinyl chloride
were also active in this test system, while bromomethane, meth
ane, and ethane were inactive. Incorporation of some of these
compounds into liquid cell culture medium for cell treatment was
either unsuccessful or produced only a weak enhancement re
sponse. Methodology is now available to evaluate volatile and
gaseous carcinogens or mutagens and can be used to identify
their mechanisms of action and the relative hazards of these
agents to human health.
INTRODUCTION
To prevent human cancers caused by exposure to environ
mental carcinogens, suitable methodologies and bioassays must
be developed that will permit the detection of all classes and
physical forms of carcinogenic agents. Quantitative cell culture
assay systems provide procedures for detecting potential carcin-
'This work was supported by the United States Environmental Protection
Agency under Contract No. 68-02-2566. A preliminary account of this work was
presented at the Symposium on Genotoxic Effects of Airborne Agents, Brookhaven,
N. Y., February 1981 (13). This paper has been reviewed by the Health Effects
Research Laboratory, United States Environmental Protection Agency, and ap
proved for publication. Mention of commercial products does not constitute en
dorsement or recommendation for use.
2To whom requests for reprints should be addressed.
Received September 16,1982; accepted January 14,1983.
MAY 1983
ogens in a relatively short period of time compared to in vivo
tests. Mammalian cells in culture are particularly appropriate for
such assays because they represent an extension of the in vivo
experimental models, eliminate host factors, afford a means to
control test chemical levels, and can be conducted in a controlled
environment. Quantitative linear dose-response relationships
have been demonstrated for cell transformation assays using
fibroblasts from hamster embryo cells (1, 2, 8, 9). The transfor
mation event data are consistent with a one-hit phenomenon
indicating a direct cause-and-effect relationship (7, 8).
A short-term in vitro assay for carcinogens and mutagens has
been developed (2,4, 5, 8,13,15) that determines the ability of
various chemical carcinogens to enhance the adenovirus trans
formation of SHE3 cells. The viral enhancement assay reflects
the capacity of a chemical to damage cellular DNA either directly
or indirectly and provides a rapid mammalian cell bioassay sys
tem for screening suspected genotoxic environmental chemicals.
This bioassay has detected activity for a large number of com
pounds from a variety of chemical classes, including alcohols
and phenols, aliphatic amines, alkyl sulfates and sultanes, aro
matic amines, aryl halides, carbohydrates and derivatives, hy-
droxylamines, mycotoxins, polycyclic hydrocarbons, hydrazines,
metal salts, steroid hormones, and chlorinated hydrocarbons (2,
3). The last 4 classes are generally negative in standard bacterial
(Salmonella) assays (21, 22). Data from approximately 105 test
performances in 29 different chemical classes have been pub
lished for the enhancement of viral transformation assay (2, 3).
With that assay as an index of carcinogenicity, 94% of 130
chemicals with known positive or negative activity agreed with
their current in vivo classifications regarding carcinogenicity.
Volatile or gaseous chlorinated compounds are ubiquitous
environmental contaminants. Their use in industrial workplaces
is widespread; for example, the annual production of 1,2-dichlo
roethane is 11.8 billion pounds/year in the United States (17).
Small amounts of chloroform and related chemicals can be
detected in municipal drinking water supplies (23), and DCM is
used in certain food extraction processes (17).
Previous attempts to evaluate the genotoxic potential of TCE
and 1,2-dichloroethane by viral enhancement assays with incor
poration into liquid cell culture media were unsuccessful or
produced only weak or inconsistent enhancement responses.
This report documents the successful development of methods
for treating mammalian cell cultures with volatile organic liquids
and gaseous compounds and measuring the chemical enhance
ment of DNA viral transformation in hamster embryo cells. The
compounds studied were: TCE, DCM, 1,2-dichloroethane, 1,1-
3 The abbreviations used are: SHE, Syrian hamster embryo; TCE, 1,1,1 -trichto-
roethane: DCM, dichloromethane; MDM, modified Dulbecco's medium; FBS. fetal
bovine serum; TF, transformation frequency.
This One
1945
DWJC-958-74GG
on May 1, 2016. © 1983 American Association for Cancer Research. cancerres.aacrjournals.org Downloaded from