Oxidation of ABTS by Silicate-Immobilized Cytochrome c in
Nonaqueous Solutions
Joseph Deere, Edmond Magner,* J. Gerard Wall, and B. Kieran Hodnett
Materials and Surface Science Institute and Department of Chemical and Environmental Sciences,
University of Limerick, Limerick, Ireland
Cytochrome c can be readily adsorbed onto mesoporous silicates at high loadings of
up to 10 mmol g
-1
of silicate. The adsorbed protein retains its peroxidative activity,
with no diffusional limitations being observed. The protein can be adsorbed onto the
external surface of the silicate or, provided that the pore diameter is sufficiently large,
into the channels. In aqueous buffer, the catalytic activity of the adsorbed protein (for
the oxidation of ABTS) decreased with increasing temperature, with the decrease being
less marked for cytochrome c held within the silicate channels. Similar results were
obtained in 95% methanol. Analysis of kinetic data showed that significant increases
in k
cat
/K
M
occurred in methanol, ethanol, and formamide, with slight decreases
occurring in 1-methoxy-2-propanol. The observed increases were primarily a result of
substantial increases in k
cat
, while the results in 1-methoxy-2-propanol can be ascribed
to increases in K
M
. Resonance Raman spectroscopy indicated that the structure of the
heme environment of the adsorbed protein was essentially unchanged, in aqueous
buffer and in the nonaqueous solvents, methanol, 1-methoxy-2-propanol, and ethanol.
In addition, Raman spectra of the lyophilized protein indicated that there were no
apparent changes in the heme structure.
Introduction
The use of silicates to immobilize proteins is well-
established (1-3). Weetall (4) has reviewed the use of
controlled pore glass (CPG) to immobilize proteins. These
studies involved the use of CPG of pore sizes ranging
from 300 to 2000 Å, with immobilization occurring via
covalent linkage of the protein or enzyme to the glass. A
wide range of proteins have been immobilized onto CPG.
In general, it is necessary that the pore size of the glass
be significantly larger than the biological molecule of
interest. The amount of protein that can be immobilized
depends on the pore size and surface area of the glass,
with maximal activity occurring with material possessing
both an optimal pore size and surface area (5). The major
disadvantages in using CPG materials are their cost and
more importantly their surface areas, which rapidly
decrease with increasing pore size (5, 6).
The use of sol-gel-derived silica glasses to encapsulate
proteins has been well-documented. The synthesis of such
glasses involves the hydrolysis of siloxanes, which upon
polymerization and condensation yield a solid (7). The
biological molecule of interest is placed in the reaction
mixture and encapsulated within the solid as it is formed.
A wide range of redox proteins, including cytochrome c
(cyt c), hemoglobin, horseradish peroxidase, glucose oxi-
dase, and nitrate reductase, have been encapsulated
within sol-gels (7). The majority of these studies have
focused on the development of sensors, utilizing the
encapsulated biological component as the recognition
element. The focus on the development of sensors is a
consequence of the slow rates of diffusion of substrates
through the glass to the binding sites of the protein or
enzyme. Since changes in the concentration of the bio-
recognition element will not adversely affect the response,
diffusion-limited responses are highly advantageous in
such sensors. However, such a response limits the use of
these materials in biocatalysis, where slow substrate
diffusion may not be desirable. A recent report has
described the use of sol-gel-encapsulated horseradish
peroxidase as a biocatalyst; however, the rate of reaction
was still limited by mass transport (9).
A range of mesoporous silicates (MPS) have been
described since the first material was reported by Beck
et al. (9). These materials are formed via surfactants such
as cetyl-trimethylammonium bromide in solution, which
act as structure directing agents. Addition of a silane
results in silica polymerization around the surfactant
micelle structures, forming a gel. This gel is set ther-
mally, condensing the silane to form a stable material.
On subsequent removal of the surfactant, a mesoporous
structure remains. MPS exhibit highly ordered pore
structures and very tight pore size distributions, as
exemplified by the first type of these materials reported,
MCM-41 (9). As a result of their mesopore structures,
MPS possess large surface areas of the order of 1000 m
2
g
-1
. Their silicate inorganic framework allows for a
chemically and mechanically stable material (10), which
is also resistant to microbial attack. In addition, it is
possible to chemically modify MPS with various func-
tional groups enabling electrostatic attraction or repul-
sion between MPS and the protein(s) of interest to be
maximized (11, 12). Unlike sol-gel immobilization, pro-
tein encapsulation in MPS occurs after synthesis of the
support.
MPS hold promise for use as supports to immobilize
enzymes (11-14) and may find applications in bio-
* To whom correspondence should be addressed. Tel: +353-61-
202629. Fax: +353-61-202568. E-mail: edmond.magner@ul.ie.
1238 Biotechnol. Prog. 2003, 19, 1238-1243
10.1021/bp0340537 CCC: $25.00 © 2003 American Chemical Society and American Institute of Chemical Engineers
Published on Web 05/22/2003