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Mar Biol
DOI 10.1007/s00227-014-2579-3
ORIGINAL PAPER
Impact of eutrophication and climate change on fish
and zoobenthos in coastal waters of the Baltic Sea
Martin Snickars · Benjamin Weigel · Erik Bonsdorff
Received: 14 April 2014 / Accepted: 10 November 2014
© Springer-Verlag Berlin Heidelberg 2014
benthic-feeding fish exploring the deep waters. An inten-
sification of these trends is likely in the future, as climate
change scenarios suggest further temperature increase and
salinity decrease.
Introduction
A large number of fish species utilize coastal waters during
one or several life stages (Seitz et al. 2013). One reason for
this is the high productivity of the benthic habitats, which
are central for trophic links, as many fish species feed on
zoobenthos in these shallow environments (Hüssy et al.
1997; Tomczak et al. 2009). However, multiple stressors
pose pressures on growing portions of marine and coastal
waters. For example, eutrophication and climate change are
recognized as two major problems that alter coastal ecosys-
tems (Fields et al. 1993; Cloern 2001; Harley et al. 2006),
making these habitats vulnerable as they encounter the
impacts of both.
The effects of eutrophication on marine ecosystems are
broad. Nutrient enrichment induces enhanced pelagic pri-
mary production, leading to decreased Secchi (photic) depth
and an elevated risk of low oxygen levels in the bottom
water. These effects have many ecosystem consequences,
affecting species across photic and aphotic habitats and
trophic levels (Cederwall and Elmgren 1990; Bonsdorff
et al. 1997; Conley et al. 2011). The response of zooben-
thos to eutrophication often is nonlinear. Initially, biomass
may increase due to higher food supply from increased
pelagic primary production, but eventually the biomass
begins to decrease, e.g., due to low oxygen levels in deep
water, as the impact of eutrophication progresses (Pearson
and Rosenberg 1978; Diaz and Rosenberg 2008). In coastal
waters (10–30 m), oxygen has normally been considered
Abstract Coastal waters are important in linking zooben-
thos and fish, as many coastal fishes feed on benthic prey
in these habitats. Major drivers, such as eutrophication and
climate change, may alter this link, whereas shifts in the
importance of these drivers may induce different responses
in zoobenthos and fish, respectively, potentially changing
productivity of coastal ecosystems. The aim of this study
was to assess in which way abundance and distribution
of benthic-feeding fish and biomass of zoobenthos have
changed over time in response to eutrophication (Sec-
chi depth) and climate change (temperature and salinity),
respectively. This was done by analyzing gross changes in
the responses over three decades (1983–2012) and across
depth zones. Eutrophication and climate change caused
different though specific impacts on fish and zoobenthos,
respectively. Throughout the 1980s, increasing benthic-
feeding fish abundance in shallow waters (<6 m) was pri-
marily attributed to eutrophication (decreasing Secchi
depth), implying increased system productivity. During the
2000s, the effect of eutrophication levelled out, whereas
temperature caused contrasting development of fish abun-
dance at different depth zones. Shallow waters had lower
fish abundances during warm years compared with colder
ones, while the abundance increased in deeper, aphotic
waters (6–20 m). In the deep waters, zoobenthos showed a
contrasting, declining trend in biomass, coinciding with the
decrease in salinity during the 2000s. This suggests altered
ecosystem productivity and potential food shortage for
Communicated by F. Bulleri.
M. Snickars (*) · B. Weigel · E. Bonsdorff
Department of Biosciences, Åbo Akademi University,
Artillerigatan 6, 20521 Åbo, Finland
e-mail: msnickars@gmail.com