FEMS Microbiology Letters69 (lge~0)145-1:52 145
Publishedby Elsevier
FEMSLE03998
Production of novel pullulanases at high concentrations
by two newly isolated thermophilic clostridia
M. Klingeberg 1, H. Hippe 2 and G. Antranikian 3
) Teefinische Universitiit ttamburg-Harburg, Hamburg, ~lnstitut flit Mikrobialagie der Unwersitiil Giittingen, ti&tingen
and ' German Collection of Microorganisms and Cell Cultures (DSM), Braunschweig F.R.G.
Receivedit Januar~1990
Revision r'ec¢ived 23 Januar'j 1090
Accepted24 January I990
Key words: CIostridium thermohydrosulfuricum; Thermophilic bacteria; Pullulanase type II;
Enzyme secretion
1. SUMMARY
Two thermophilic bacteria, which are capable
of vowing on starch at 60-70°C under anaerobic
conditions, were isolated from a sugar refinery in
Uelzen and from Solar lake in Israel. On the basis
of their physiological characteristics they were
identified as Clostridium thermohydrosulfuricum
Uel 1 and C. thermohydrosulfuricum Sol 1, respec-
tively. The product pattern of glucose polymer
hydrolysis showed that both strains secreted en-
zymes that possess amylolytic and pullulytic activ-
ities. The major product formed was maltose. In
addition, ,t.ghicosidase activity could be detected
in the supernatams of Uel 1 strain. Compared to
most anaerobes investigated these isolates secreted
extremely high concentrations of pulhilanases in
batch culture. Up to g57~ of the total enzyme
synthesized was detected in the culture fluid. Un-
like the pullulanases of type I, which can only
Correspondence 1o: Prof. Dr. G. Anlrar~tdan. lastilut fiir
Mikrobiologie tier Georg-August Univcrsit/it Gi~ttingen. Grlm-
bachstrass¢ 8. 3400Ge, tting~nF.R.G,
attack the a-l,6-glycosidic linkages, the pul-
lulanases of both dostridial strains were also ca-
pable of hydrolyzing a-l,4-1inkages. The enzyme
system of both bacteria was found to be highly
thermoactive; optimal activity was detected at pH
5,0 and 85°C. Even at 950C and without the
addition of metal ions still 15% to 25% of en-
zymatic activity was detectable.
2. INTRODUCTION
Aerobic as well as anaerobic bacteria have de-
veloped various strategies for the utilization of
starch 11,2], a highly branched polysaecharide
which is composed of glucose units bound by
a-lA- and a-l,6-glycosidie linkages. In the last
decade a number of thermophilic anaerobic
eubacteria were isolated. Most of them belong to
the genera Clostridium, Thermoanaerobacter,
Thermobacteroides and Dictyogtomus. The major-
ity of these organisms synthesize a-amylase as
well as puilulanase. The formation of pullulanase,
which is capable of hydrolyzing a-l,6-glycosidic
linkages in pullulan in addition to a-l,4-1inkages
0378-1097/90/$03.50© 1990Federation of EuropeanMicrobiological Societies