4 A Review on the Role of
Endozoochory in Seed
Germination
A. TRAVESET , A.W. ROBERTSON AND J. RODRÍGUEZ-PÉREZ
Introduction
A large fraction of living plants produce fruits that attract animals (64% and
27% of gymnosperm and angiosperm lineages, respectively; Herrera and
Pellmyr, 2002). In consuming them, animals can spread the seeds to more
or less distant sites from the parent plant, thus contributing to plant
regeneration and colonization of new sites (Willson and Traveset, 2000, and
references therein). For millions of years both the pulp and seeds of fruit
have been subjected to selective pressures exerted by frugivores and
granivores. Fruit consumers, specifically, may show preferences for fruit
traits such as size, shape, chemical composition and others, and have
specific morphologies and physiologies of the digestive tract that affect the
survival probability of the ingested seeds in different ways (Herrera and
Pellmyr, 2002, and references therein). Seed traits are under further
selective pressures imposed by a number of biotic (antagonistic fungi,
insects and microbes; Cipollini and Levey, 1997) and abiotic factors (e.g.
light, temperature, rainfall; Holl, 1999; Leishman et al., 2000) that influence
the dormancy period, germination time and/or future seedling growth,
which can ultimately determine germination and seedling success (Verdú
and Traveset, 2005, and references therein). Therefore, the specific seed
traits in an endozoochorous plant are a result of the combined selection
imposed by frugivores, granivores and other biotic and abiotic factors.
In this chapter, we evaluate the role of endozoochory on seed
germination. We review the information gathered on germination patterns
in experiments aimed at examining the effect of a seed’s passage through a
frugivore’s gut, and give further directions on methods for future studies.
Fruit treatment in the disperser’s digestive tract (which includes pulp
separation from seeds and treatment of the seeds) can determine the
capacity of seeds to germinate and, thus, is one of the components of
78 © CAB International 2007. Seed Dispersal: Theory and its Application in a
Changing World (eds A.J. Dennis et al.)
04Seed Dispersal Ch 4 16/4/07 15:53 Page 78