Breast Cancer Research and Treatment 83: 221–231, 2004.
© 2004 Kluwer Academic Publishers. Printed in the Netherlands.
Report
Growth inhibitory activity of extracts and purified components
of black cohosh on human breast cancer cells
Linda Saxe Einbond
1
, Masahito Shimizu
1
, Danhua Xiao
1
, Paiboon Nuntanakorn
2
, Jin T. E.
Lim
1
, Masumi Suzui
1
, Colette Seter
1
, Thomas Pertel
1
, Edward J. Kennelly
2
, Fredi
Kronenberg
1
, and I. Bernard Weinstein
1
1
Columbia University College of Physicians and Surgeons, New York, NY;
2
Lehman College in the Graduate
Center, The City University of New York, Bronx, NY, USA
Key words: Actaea racemosa, actein, Cimicifuga racemosa, cinnamic acid ester, cyclin D1, triterpene glycoside
Summary
The purpose of this study was to determine whether black cohosh contains constituents that inhibit the growth of
human breast cancer cells, and therefore might eventually be useful in the prevention or treatment of breast cancer.
Black cohosh rhizomes were extracted with methanol/water and fractionated by solvent–solvent partitioning to
yield three fractions: hexane, ethyl acetate and water. The ethyl acetate fraction displayed the highest potency
in two cell-based assays, growth inhibition and cell cycle analysis. This fraction inhibited growth of both the
ER
+
MCF7 and ER
-
MDA-MB-453 human breast cancer cell lines with IC
50
values of about 20 and 10 μg/ml,
respectively. It also induced cell cycle arrest at G1 when tested at 30 μg/ml and at G2/M at 60 μg/ml in MCF7 cells.
This suggests that the extract contains a mixture of components with the more active (or more abundant) causing
G1 arrest and the less active causing G2/M arrest. We then examined specific components in this extract. The
triterpene glycoside fraction obtained by polyamide column chromatography, and the specific triterpene glycosides
actein, 23-epi-26-deoxyactein and cimiracemoside A, inhibited growth of the MCF7 human breast cancer cells and
induced cell cycle arrest at G1. The most potent compound, actein, decreased the level of cyclin D1, cdk4 and
the hyperphosphorylated form of the pRb protein and increased the level of p21
cip1
in MCF7 cells, changes that
may contribute to the arrest in G1. Further studies are in progress to identify the mechanisms by which actein and
related compounds present in black cohosh inhibit growth of human breast cancer cells.
Introduction
There is growing interest in the use of herbs to aid in
the maintenance of women’s health. Plants contain a
wide variety of chemicals that have potent biological
effects, including anticancer activity [1]. Identification
of the active components, their mechanisms of action,
and their possible interactions is important in order
to assess their potential for clinical use and possible
adverse side effects.
Black cohosh (Actaea racemosa L. syn. Cimici-
fuga racemosa {L.} Nutt) is a North American pe-
rennial in the buttercup family (Ranunculaceae) that
has been in use in Europe for the last 50 years as
a natural alternative to hormone replacement therapy
and for other gynecological conditions [2]. The plant
grows in the understory of hardwood forests east of
the Mississippi River, especially in Ohio and West
Virginia [3]. Related Asian species have been em-
ployed in Traditional Chinese Medicine to treat infec-
tious diseases [4]. The rhizome has been in use for
hundreds of years by Native Americans for a variety of
purposes that include inflammatory conditions, stimu-
lation of menstrual flow, dysmenorrhea, suppression
of cough, treatment of diarrhea, and rheumatism [5].
American women are increasingly turning to black
cohosh as a ‘more natural’ alternative to estrogen re-
placement therapy, in the belief that it has the benefits,