1 3
Mar Biol
DOI 10.1007/s00227-015-2617-9
SHORT NOTE
Species-specific differences in thermal tolerance may define
susceptibility to intracellular acidosis in reef corals
Emma M. Gibbin · Hollie M. Putnam · Ruth D. Gates ·
Matthew R. Nitschke · Simon K. Davy
Received: 29 August 2014 / Accepted: 16 January 2015
© Springer-Verlag Berlin Heidelberg 2015
placed them in a live-cell chamber attached to a confocal
microscope and bathed them in CO
2
-acidified seawater
(~pH 7.6) for 30 min, before measuring the light-adapted
pH
i
of both the host cell and its symbiont. Cells isolated
from P. damicornis were more prone to cellular acido-
sis (declines in pH
i
of 11 and 8 % in host and symbiont,
respectively, at 31 °C relative to 23.8 °C) than cells isolated
from M. capitata (5 and 4 %, respectively). These results
highlight the important role of Symbiodinium productivity
(in addition to a range of physico-chemical factors such as
skeletal morphology and tissue pigmentation) in determin-
ing the sensitivity of corals to rising sea surface tempera-
tures and ocean acidification.
Introduction
Post-industrial atmospheric CO
2
levels are increasing at a
rate and magnitude that is unparalleled in recent geological
history. To date this has resulted in a 1–2 °C increase in the
global mean temperature and a 0.08 reduction in oceanic
surface water pH, from 8.13 to the present day 8.05 (IPCC
2014). The combination of increasing temperature, declin-
ing pH and a reduction in the amount of available car-
bonate via ocean acidification is predicted to have severe
consequences for calcifying species, potentially leading to
phase shifts towards ecosystems dominated by algal- or
soft-bodied organisms (Bell et al. 2013). Coral reefs are
already showing signs of demise, particularly in areas that
are subjected to sustained periods of exposure to warm
water (Carpenter et al. 2008). Chronic stress often results
in the photosynthetic dysfunction and subsequent loss of
single-celled symbiotic dinoflagellates (Symbiodinium sp.)
and/or their photosynthetic pigments (Weis 2008). This
collapse in symbiosis, a process termed “coral bleaching”,
Abstract It is widely acknowledged that temperature
stress affects an organism’s sensitivity to ocean acidifi-
cation and vice versa, yet it is not clear how the two are
mechanistically linked. Here, we induced thermal stress in
two coral species with differing bleaching susceptibilities
to measure how a reduction in photosynthetic performance
impacts intracellular pH (pH
i
) regulation in the symbiotic
dinoflagellates (Symbiodinium sp.) and their host coral
cells. Our hypothesis was that thermally induced photo-
synthetic dysfunction in the symbiont would prevent the
efficient removal of additional CO
2
, lowering its buffering
capacity and thus increasing the host cell’s susceptibility to
intracellular acidosis. To test this, we exposed Pocillopora
damicornis (a thermally sensitive coral) and Montipora
capitata (a thermally resilient coral) to four different tem-
perature treatments (23.8, 25.5, 28 and 31 °C) for 1 week.
We then isolated intact symbiotic coral endodermal cells,
Communicated by R. Hill.
Electronic supplementary material The online version of this
article (doi:10.1007/s00227-015-2617-9) contains supplementary
material, which is available to authorized users.
E. M. Gibbin · S. K. Davy (*)
School of Biological Sciences, Victoria University of Wellington,
Kelburn Parade, Wellington 6140, New Zealand
e-mail: simon.davy@vuw.ac.nz
E. M. Gibbin
e-mail: emma.gibbin@hotmail.co.uk
H. M. Putnam · R. D. Gates
Hawaii Institute of Marine Biology, University of Hawaii,
PO Box 1346, Kaneohe, HI 96744, USA
M. R. Nitschke
School of Biological Sciences, The University of Queensland,
St Lucia, Brisbane, QLD 4072, Australia