NEttnOSCt [
LETTERS
ELSEVIER Neuroscience Letters 168 (1994) 213 216
Alteration of pallidal cholinergic activity in MPTP-treated monkeys:
effect of dihydro-a-ergocryptine (DEK)
D. Curti*, E. Izzo, G. Benzi
Institute of Pharmacology, Faculty of Sciences, University of Pavia, 27100 Pavia, Italy
Received 14 April 1993; Revised version received 20 December 1993: Accepted 23 December 1993
Abstract
Monkeys, intravenously administered with MPTP at the dose of 0.3 mg/kg for 5 consecutive days, develop a severe Parkinson-like
syndrome. Cholinergic enzyme activities are increased in the internal segment of the globns pallidus (GPi) and into a lesser extent
in the external globus pallidus (GPe). Cholinergic activities are not significantly affected in the caudate and putamen nor in the
frontal, parietotemporal, occipital cortices and in the cerebellum. The treatment of the animals twice daily for 2 weeks with
dihydro-c~-ergocryptine (DEK) starting 5 days before the first MPTP administration counteracts the neurotoxin-induced alteration
in the internal pallidum and ameliorates some motor related parkinsonian symptoms.
Key words: Monkey; MPTP; Parkinsonism; Cholinergic activity; Dihydro-c~-ergocryptine; Globus pallidus
The neurotoxin 1-methyl-4-phenyl-l,2,3,6-tetrahy-
dropyridine (MPTP) induces Parkinson (PD)-like symp-
toms in human and non-human primates [4,12]. MPTP
is converted by the monoamino oxidase type B (MAO-B)
to MPP + (1-methyl-4-phenylpyridine) which is taken up
by the dopamine (DA) carrier, accumulates intraneu-
ronally and rather selectively destroys the dopaminergic
neurons in the pars compacta of the substantia nigra
(SNc) causing a marked decrease of the DA content in
the caudate and putamen nuclei [4]. The neostriatum is
the site where the DA-acetylcholine (Ach) interaction
are reported to occur principally. The prevailing opinion
(although there is no general agreement [6]) is that the
DA released from nigro-striatal terminals tonically in-
hibits, through stimulation of O 2 receptors, the release
of Ach from striatal cholinergic interneurons [15,27]. D 2
receptors antagonists remove this inhibitory modulation
and increase Ach release in the striatum [28]. Central
dopaminergic and cholinergic systems seem to exert a
mutual inverse relationship [1]; however, the effect of
nigrostriatal denervation on cholinergic activity outside
*Corresponding author. Fax: (39) 382-386 385.
0304-3940/94157.00 © 1994 Elsevier Science Ireland Ltd. All rights reserved
SSDI 0304-3940(94)00008-X
the neostriatum is not understood and it is likely to play
a role in Parkinson's-associated motor disturbances; in
fact cholinergic stimulation of GPi elicits rigidity of the
paravertebral musculature and increases tremor ampli-
tude in PD patients [5].
The aim of this investigation was to evaluate in some
striatal and extrastriatal brain regions the short term
consequences of MPTP administration on the choliner-
gic enzymes (choline acetyltransferase, CHAT, and ace-
tylcholinesterase, AchE) and on pyruvate dehydrogenase
complex (PDHc) which catalyzes the oxidative decar-
boxylation of pyruvate to acetylcoenzyme A, linking
cholinergic and energy metabolism. A treatment with
dihydro-a-ergocryptine (DEK), a D2 receptor agonist,
was also performed. This ergot derivative improves
motor performances (rigidity, bradykinesia and tremor)
in PD patients [21].
Male cynomolgus monkeys were housed at the primate
facility of the Istituto di Ricerche Biomediche 'Antoine
Marxer' RBM (Ivrea, Italy). Animal care and handling
were in accordance with fundamental principles by
CIOMS and with provisions of EEC Council Directive
86-609. The animals were randomly divided into three
groups: MPTP-treated animals (n = 6) received a dose of