1 Type of article: Original article, second submission. Original MS no. 12672 1 Title: Effect of light conditions on anatomical and biochemical aspects of somatic and 2 zygotic embryos of hybrid larch (Larix x marschlinsii) 3 Authors: Patrick von Aderkas 1* , Caroline Teyssier 2 , Jean-Paul Charpentier 2 , Markus 4 Gutmann 3 , Luc Pâques 2 , Claire Le Metté 2 , Kevin Ader 2 , Philippe Label 4 , Lisheng Kong 1 , 5 Marie-Anne Lelu-Walter 2 6 Affiliations: 7 1 Centre for Forest Biology, Department of Biology, University of Victoria, 3800 Finnerty 8 Rd., Victoria BC, V8W 3N5, Canada, Fax: + 12507216611, E-mail: pvonader@uvic.ca 9 2 I.N.R.A., UR 0588 Research Unit on Breeding, Genetic and Physiology of Forest Trees, 10 2163 Avenue de la Pomme de pin CS 4001 Ardon F- 45075 Orléans Cedex 2 France 11 3 Fachgebiet Obstbau, Technische Universität München, Dürnast 2, D-85354 Freising, 12 Germany 13 4 INRA-Université Blaise Pascal, UMR_A 547 PIAF, Les Cézeaux, 24 Avenue des 14 Landais, 63177 Aubière cedex, France. 15 Running title: larch somatic and zygotic embryogenesis and the effect of light 16 Email addresses 17 Patrick von Aderkas – pvonader@uvic.ca 18 19