Journal of Neuroscience Methods, 10 (1984) 83-101 83
Elsevier
NSM 00361
Research Papers
A method for rostrocaudal integration of
morphometric information from
transmitter-identified cell groups. A morphometrical
identification and description of 5-HT cell groups in
the medulla oblongata of the rat *
L.F. Agnati 1, K. Fuxe 2 L. Calza 1, I. Zini 1 T. H6kfelt 2, A. Steinbusch 2
and A. Verhofstad 3
I Departments of Human PhysioloD' and Endocrinology, Unwersity of Modena, Modena, (Italy),
2 Department of Histology, Karolinska Institutet, Stockholm (Sweden) and 3 Department of Anatomy and
Embryology, Catholic University, Nijmegen (The Netherlands)
(Received February 4th, 1983)
(Revised September 28th, 1983)
(Accepted December 23rd, 1983)
Key words: morphology--5-HT--morphometry--immunocytochemistry--cell body parameters--cell
group parameters--rostro-caudal integration--medulla oblongata
A method has been developed to integrate rostrocaudal information from morphometrically char-
acterized 5-HT nerve cell groups visualized by means of the indirect immunofluorescence technique in
single coronal sections of the neuroaxis. The present method has been applied to the 5-HT positive cells of
the medulla oblongata of the rat. 5-HT cell body and cell group parameters were measured by the use of a
semiautomatic image analyzer (Kontron, MOP AMO 2) plugged into an Apple II computer.
The density distribution of 5-HT positive cells was also studied by dividing the area analyzed into
unitary squares (about 120× 120 p,m) and then by considering the number of 5-HT cells falling in each of
these squares. The existence of a 5-HT cell group was determined by testing the randomness of the 5-HT
profiles per unitary square. The entire population of 5-HT positive cells as well as the 5-HT cell groups
were described in terms of gravity center coordinates, mean maximal diameter and homogeneity index. A
rostrocaudal representation of the gravity centers and of the dispersion of the 5-HT nerve cell groups
around them provided an exact three-dimensional description of the respective locations of the 5-HT cell
groups within the medulla oblongata. These methods can be applied to all types of transmitter-identified
cell groups and at any level of the neuroaxis.
* In memory of Professor Giorgio Montanari, professor at the Medical Faculty of Modena.
Correspondence." K. Fuxe, Department of Histology, Karolinska Institutet, Box 60400, S-104 01 Stock-
holm, Sweden.
0165-0270/84/$03.00 © 1984 Elsevier Science Publishers B.V.