ELSEVIER Mutation Research 305 (1994) 145-150
Fundamentaland Molecurar
Mechanisms of Mutagenesis
The anticlastogenic effect of tocopherol in peritoneal
macrophages of benznidazole-treated and ovariectomized mice
Zulmira G.M. Lacava *, Helena Luna
Department of Genetics and Morphology, Institute of Biological Sciences, Universityof Brasilia, 70910-900 Brasilia, DF, Brazil
(Received 12 January 1993)
(Revision received 8 August 1993)
(Accepted 1 September 1993)
Abstract
Cytogenetic studies revealed a significant increase in the frequency of structural chromosome aberrations of
peritoneal macrophages from hyperimmune Swiss mice after ovariectomy. The administration of the nitroarene
benznidazole caused a large number of chromosomal deletions in peritoneal macrophages of sham-ovariectomized
animals. The clastogenic effect of benznidazole was much greater in peritoneal macrophages of ovariectomized mice.
The anti-oxidant a-tocopherol protected the peritoneal macrophages from developing ovariectomy- or
benznidazole-induced chromosomal aberrations, thus suggesting free radical damage in these processes.
Key words: Benznidazole; Cytogenetic damage; Anticlastogenic eftect; Macrophage; Ovariectomy
1. Introduction
Benznidazole (BZ) is a nitroarene compound
used in the treatment of Chagas' disease
(American trypanosomiasis)caused by Try-
panosoma cruzi. It has been shown that BZ is
mutagenic in bacteria (Nagel and Nepomnaschy,
1983; Ohnishi et al., 1983) and clastogenic in T.
cruzi (Gouman et al., 1985). Induction of chromo-
somal aberrations and micronuelei and an in-
crease in the frequency of expression of fragile
sites were reported in cultures of peripheral blood
lymphocytes from chagasic children treated with
* Corresponding author.
BZ (Gorla et al., 1988; Moya and Trombotto,
1988) or in bone marrow cells of mice treated
with the drug (Navarro et al., 1984).
Although the mechanisms of BZ induction of
chromosomal aberrations have not been deter-
mined, it has been shown that this trypanocidal
nitroarene undergoes enzymatic reduction of the
nitro group and generates free radicals such as
nitro anions, hydroxyl, singlet oxygen and hydro-
gen peroxide (Docampo et al., 1971; Moreno et
al., 1982). The free radicals are known to cause
DNA alterations such as base damage, single-
and double-strand breaks, cross-linking, chromo-
somal aberrations (Sun, 1990) and peroxidative
damage to membrane lipids (Chow, 1991). How-
ever, the deleterious action of some free radicals
might be prevented by anti-oxidants. In this con-
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