Physiology&Behavior, Vol. 55, pp. 35-38, 1994 0031-9384/94 $6.00 + .00
Printed in the USA. Copyright© 1993PergamonPressLtd.
Estrogen Antibodies Reduce Vulnerability to
Stress-Induced Failure of Intrauterine
Implantation in Inseminated Mice
DENYS DECATANZARO, t ELAINE MACNIVEN, TINA GOODISON AND DAVID RICHARDSON
Department of Psychology, McMaster University, Hamilton, Ontario, L8S 4K1 Canada
Received 13 January 1993
DECATANZARO, D., E. MACNIVEN, T. GOODISON AND D. RICHARDSON. Estrogen antibodies reduce vulnerability to
stress-induced failure of intrauterine implantation in inseminated mice. PHYSIOL BEHAV 55(1) 35-38, 1994.--In Experiment
1, inseminated mice were randomly assigned to either an undisturbed control condition or four conditions involving exposure
to restraint stress on days l through 5 of pregnancy. Restrained animals received one of three doses of estrogen antibodies or just
vehicle injections on each day of restraint. Restrained animals receiving vehicle only showed significantly fewer uterine implantation
sites than did unrestrained controls, while restrained animals given estrogen antibodies showed more implantation sites than did
vehicle-treated restrained animals. In Experiment 2, varied dosages of refined estrogen antibodies were administered to inseminated
females concurrent to restraint-stress on days 1 through 5 of pregnancy. More females receiving the higher dosages of antibodies
produced litters than did restrained females with just vehicle injections, and at the highest dose the number of litters was similar
to that produced by undisturbed control females. These results converge with other evidence to suggest that stress-induced
pregnancy blocks are mediated by estrogens.
Stress Pregnancy disruption Implantation Estrogen Estrogen antibodies Restraint
A variety of psychological stressors have adverse effects upon
early pregnancy in mammals (5). Pregnancy can be blocked by
diverse stressors applied during the preimplantation period, in-
cluding chronic restraint (8,1 l, 17), daily handling (l 4), over-
crowding (2), forced swimming or loud noise (19), high ambient
temperatures (16), exposure to strange males (1), and predator
exposure (3,11).
The hormonal mechanisms underlying such pregnancy blocks
need further definition. Existing data suggest that hormones
normally associated with stress, adrenal corticosteroids, and cat-
echolamines, may not be as important in the mediation of this
effect as are the primary gonadal hormones, progesterone, and
estrogen (4,6, l 1). Indeed, relatively minute amounts of exoge-
nous estrogens will reliably block pregnancy, whereas exogenous
corticosterone is apparently without effect even at high dosages
(6). A previous study examined the effects of chronic restraint
stress on steroid levels in early pregnancy in rats, indicating that
endogenous estradiol levels were enhanced in stressed animals
as compared to controls (12).
If pregnancy blocks are caused by excessive estrogen levels
induced by stress, then the administration of an agent effective
in decreasing estrogen activity might allow pregnancy to continue
in the face of stress. Biological actions of a hormone can be
neutralized or diminished through the introduction of a high-
Requests for reprints should be addressed to Denys deCatanzaro.
affinity antibody via passive immunization (15). The purpose
of these experiments was to investigate whether a stress-induced
pregnancy block could be counteracted through the administra-
tion of an estrogen antibody.
EXPERIMENT 1
This experiment was conducted using serum extracted from
rabbits treated or not treated with estradiol, and uterine im-
plantation sites were examined rather than allowing females to
bear litters.
Method
Subjects were female HS (heterogeneous strain) mice bred
in our laboratory from stock originally obtained from the De-
partment of Zoology at the University of Toronto. After weaning
and prior to insemination, females were housed in groups of
four or five in standard polypropylene cages measuring 28 x 16
X 11 (height) cm with wire grid tops allowing continuous access
to food and water. The colony room was maintained under a
reversed 10:14 h dark:light cycle and at 21 _ I°C. HS males
were housed individually under these same conditions.
When the females were at least 65 days of age, they were
each housed alone at the commencement of the dark phase of
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