Effect of Preservation Conditions on Autonomic Transmission in Rat
Small Bowel
M.O. Taha, M.M. Fraga, D.J. Fagundes, C.P. Bandeira, A. Caricati-Neto, and A. Jurkiewicz
I
N small bowel, acetylcholine (ACh) released from intra-
mural cholinergic nerves acts at nicotinic cholinergic
receptors to cause excitation of enteric nerves and at
muscarinic cholinergic receptors to cause contractions of
intestinal smooth muscle.
1
Thus, contractile function of
small bowel regulated by ACh is highly dependent on
structural and functional integrity of enteric nerves.
2
His-
tological studies performed on small bowel submitted to
cold ischemic preservation prior to transplantation suggest
that neurological lesions following prolonged ischemia se-
verely compromises contractile functions of the intestine.
3–5
However, autonomic dysfunction has not been studied in
small bowel grafts preserved for transplantation.
In order to investigate autonomic transmission in small
bowel grafts preserved for transplantation, the contractile
effects mediated by muscarinic and nicotinic cholinergic
receptors in longitudinal muscle of rat jejunum submitted
to cold ischemic preservation (Ringer-Lactate solution at
4°C) for long-term (12, 18, and 24 hours) were studied by
means of stimulation with electrical or cholinergic agonists
(carbachol and nicotine).
METHODS
Animals
The experiments were conducted with female Wistar rats, weighing
200 to 240 g (12 to 16 weeks old). The day-night cycle was constant,
with 12 hours of light and dark. The animals had free access to tap
water and pelletted chow. The animals were deprived of food for 12
hours before the experiment but were allowed water ad libitum.
The study design was approved by the Animal Ethics Committee of
the UNIFESP.
Biological Preparation
The rats were killed by sulfuric ether inhalation for removal of
small bowel. The jejunum was isolated, dissected free of surround-
ing tissues, and intraluminally washed. Jejunal segments (2 cm)
were preserved in Ringer’s lactate solution at 4°C for 12 (P12), 18
(P18), or 24 hours (P24). Jejunum segments after preservation for
12 (n = 8), 18 (n = 8) or 24 (n = 8) hours were mounted, in parallel
with freshly harvested segments (n = 24), in an isolated organ bath,
containing 10 mL of nutrient solution with the following composi-
tion (mmol/L): NaCl, 138; KCl, 5.7; CaCl
2
, 1.8; NaH
2
PO
4
, 0.36;
NaHCO
3
, 15; and dextrose 5.5 (37°C, 7.4 and bubbled with air).
Longitudinal muscle contractions induced by electrical or drug
stimulation were recorded by means of isometric transducers HP
FTA10 coupled to a HP 7754A polygraph. Electrical stimulation
(ES) was applied by means of platinum electrodes coupled to an
electrical stimulator Grass S88 (10 to 30 Hz, 1 ms, 60 to 80 V).
Jejunal segments were also stimulated by the cholinergic agonists
carbachol (CCh) or nicotine (NIC) and by a purinergic agonist
(ATP). Effects of ES and agonists were studied in absence or
presence of the cholinergic antagonists atropine (ATR; 10
-6
mol/L) or hexametonium (HEX; 10
-5
mol/L). The relaxing effects
of adrenaline (ADR; 10
-3
mol/L) were studied in preserved and
control segments. Maximal effects (E
max
) produced by ES or drugs
were determined in preserved and control segments. Statistical
analyses of E
max
were performed by analysis of variance.
RESULTS
In preserved and control jejunum, contractile response
induced by ES with 10, 20, and 30 Hz were biphasic and
composed of an initial rapid component followed by a
sustained component (Fig 1). In relation to the control, the
response induced by ES was significantly lower in segments
preserved for 12, 18, 24 hours (Fig 1; Table 1). These
contractions were blocked by ATR (10
-6
mol/L) in control
and in preserved preparations (Fig 2).
Contractions induced by CCh (10
-3
mol/L) or NIC (10
-3
mol/L) were lower in segments preserved for 12, 18, or 24
hours than in control segments (Fig 3 and 4; Table 1).
Contractions induced by CCh were blocked by ATR (10
-6
mol/L; (Fig 5) while contractions induced by NIC were
blocked by HEX (10
-6
mol/L, not shown). The relaxing
effects of ADR (10
-3
mol/L) were not different in pre-
served and control segments (Fig 5).
From the Departments of Surgery (M.O.T., M.M.F., D.J.F.,
C.P.B) and Pharmacology A.C.N., A.J.), Universidade Federal de
Sa ˜ o Paulo (UNIFESP), Escola Paulista de Medicina,Sa ˜ o Paulo-
SP, Brazil.
This was supported by Fundac ¸a ˜ o de Amparo a ` Pesquisa do
Estado de Sa ˜ o Paulo (FAPESP) and Centro de Estudos da
Disciplina de Te ´ cnica Operato ´ ria e Cirurgia Experimental
(CEDITEC)—UNIFESP/EPM.
Address reprint requests to Murched O. Taha, PhD, UNIFESP/
EPM, Te ´ cnica Operato ´ ria e Cirurgia Experimental, Rua Botucatu,
740, CEP 04023-900, Sa ˜ o Paulo-SP, Brazil.
© 2002 by Elsevier Science Inc. 0041-1345/02/$–see front matter
655 Avenue of the Americas, New York, NY 10010 PII S0041-1345(02)02698-2
Transplantation Proceedings, 34, 1021–1024 (2002) 1021