Effect of Allopurinol on Autonomic Dysfunction in Rat Jejunal
Segments Exposed to Cold Ischemic Preservation for Transplantation
M.O. Taha, M.M. Fraga, D.J. Fagundes, A. Jurkiewicz, and A. Caricati-Neto
T
HE CONTRACTILE FUNCTION of the small bowel
is mainly regulated by acetylcholine (ACh) released
from enteric cholinergic nerves, which acts at nicotinic
cholinergic receptors to stimulate ACh release and at
muscarinic cholinergic receptors to induce smooth muscle
contraction.
1
These regulatory actions of ACh are highly
dependent on the structural and functional integrity of
enteric cholinergic nerves.
2
Histological studies of small bowel after cold ischemic
preservation (CIP) for organ transplantation have shown
severe lesions of the enteric autonomic nerves.
3–5
Func-
tional studies have revealed a dramatic reduction in intes-
tinal contractile function mediated by these nerves.
6
These
autonomic dysfunctions that reduce contractile functions of
small bowel exposed to CIP apear to result from the actions
of cytotoxic substances generated under ischemic condi-
tions.
7
Ischemia injury is one of the leading causes of loss of
transplanted organs. Oxidative damage is thought to play an
important role in ischemia injury, including the outcome of
organ transplantation. Xanthine oxidase has been widely
reported to be an important source of cytotoxic free radicals
under ischemic conditions.
7
Various pharmacological
agents have been evaluated. To reduce free radical levels
under ischemic conditions.
8 –10
Some studies suggest that
allopurinol, an analogue of hypoxanthine, is a specific
potent inhibitor because it acts as a substrate for xanthine
oxidase, thereby reducing free radical generation under
ischemic conditions.
8 –12
To evaluate the long-term effects
of allopurinol on autonomic dysfunction in small bowel
exposed to CIP, for we studied the contractile functions
mediated by enteric cholinergic nerves in rat jejunal seg-
ments exposed for 12, 18, or 24 hours to CIP (4°C) in
Ringer’s lactate solution containing 0.5 mmol/L, allopuri-
nol.
METHODS
Animals
Male Wistar rats weighting 240 to 300 g (16 to 20 weeks old)
obtained from animal care UNIFESP/EPM were maintained with
day–night (12-hour light/dark) cycles and given free access to tap
water and pellet chow. The animals were deprived of food for 12
hours before the experiment but were allowed water ad lib. The
study design was approved by the Animal Ethics Committee of the
UNIFESP-EPM.
Biological Preparation
Rats sacrificed sulfuric ether inhalation underwent midline lapa-
ratomy for surgical removal of the small bowel. The jejunum was
isolated, washed intraluminally, and cleaned of surrounding tis-
sues.
13
Jejunal segments (2 cm) were exposed to CIP (4°C in
Ringer’s lactate solution) without (-) or with (+) addition of 0.5
mmol/L
-1
allopurinol (AL). After preservation for 12, 18, or 24
hours, AL- and AL+ jejunal segments were mounted in parallel
under 1 g tension in isolated organ baths containing 10 mL of
aerated nutrient solution containing (in mmol/L): NaCl 138, KCl
5.7, CaCl
2
1.8, NaH
2
PO
4
0.36, NaHCO
3
15, and dextrose 5.5 (37°C,
pH 7.4).
13
Longitudinal muscle contractions were induced by
stimulation of the enteric autonomic nerves (electrical transmural
stimulation, ETS) or nicotinic (nicotine, NIC) or muscarinic (Car-
bachol, CCh) cholinergic receptors. For ETS, jejunal segments
were mounted between two parallel platinum electrodes connected
to an electrical stimulator (Grass S88) for delivery of electrical
pulses (10 to 30 Hz, 1 ms duration and 60 V). The effects of
nicotinic (hexametonium, HEX) and muscarinic (atropine, ATR)
antagonists were studied on these contractions. Contractile re-
sponses were recorded by means of force-displacement transducers
connected via a bridge amplifier to a computerized recording
system (PowerLab, ADInstruments, USA). The maximal contrac-
tions (C
max
) produced by ETS, NIC, and CCh in AL- and AL+
jejunal segments were analyzed by and one-way analysis of variance
(ANOVA) and Student t test. A P value of .05 was considered
significant.
From the Departments of Surgery (M.O.T., M.M.F., D.J.F.) and
Pharmacology (A.J., A.C.N.), Universidade Federal de Sa ˜ o Paulo
(UNIFESP), Escola Paulista de Medicina (EPM), Sa ˜ o Paulo,
Brazil.
This work was supported by grants from the Fundo de Auxilio
dos Docentes e Alunos (FADA)—UNIFESP/EPM, Centro de
Estudos da Disciplina de Te ´ cnica Operato ´ ria e Cirurgia Experi-
mental (CEDITEC)—UNIFESP-EPM, Conselho Nacional de Des-
envolvimento Cientifico e Tecnolo ´ gico (CNPq), and Fundo de
Amparo a ` Pesquisa do Estado de Sa ˜ o Paulo (FAPESP).
Address reprint requests to Murched Omar Taha, PhD, Sur-
gery Department, UNIFESP/EPM, Rua Botucatu, 740, CEP
04023-900, Sa ˜ o Paulo-SP, Brazil. E-mail: taha@uol.com.br
© 2004 by Elsevier Inc. All rights reserved. 0041-1345/04/$–see front matter
360 Park Avenue South, New York, NY 10010-1710 doi:10.1016/j.transproceed.2004.01.095
Transplantation Proceedings, 36, 293–295 (2004) 293