Pectinases are a group of enzymes responsible for
plant tissue maceration and consist of a number of
enzymes that are classified according to their mode of
action and the substrates they prefer, viz. pectin esterases
(EC 3.1.1.11), polygalacturonases (EC 3.2.1.15), pectate
lyases (EC 4.2.2.2), and pectin lyases (PNL) (EC
4.2.2.10). Amongst these, PNL are of particular interest
because they degrade pectin polymers directly by the β-
elimination mechanism that results in the formation of
4,5-unsaturated oligogalacturonides, while other pectinas-
es act sequentially to degrade pectin molecule totally. The
major source of PNL on industrial scale is fungi belonging
to the genera Aspergillus, Penicillium, and Fusarium,
although a few bacterial PNL are also reported [1]. Solid-
state fermentation (SSF) has emerged as preferred tech-
nology in industrial fermentation as it offers several practi-
cal and economical advantages [2]. Agro-industrial
residues and wastes such as wheat bran, rice bran, sugar-
cane bagasse, corncobs, citrus wastes, apple pomace, and
a variety of such byproducts are potentially good substrates
for SSF [3]. Acidic PNL have biotechnological potential
in fruit juice industries due to the fact, that they degrade
pectin without disturbing the ester group, which is respon-
sible for specific aroma of the juice, and also it does not
lead to formation of the toxic alcohol methanol.
Though there is a considerable amount of literature
on application of acidic PNL in juice clarifying activity
[4-6], reports on retting of natural fiber by alkaline PNL
are few [7]. Retting is a process of fiber separation from
non-fiber parts of plants. The plant fibers are held togeth-
er by pectin, and pectin plays an important role as a
binder between adjacent cells [8]. Since PNL degrades
pectin, it is expected that it should play a role in the ret-
ting of natural fibers. Fibers present in the stems of many
herbaceous dicots, such as flax (Linum usitatissimum),
Indian hemp (Cannabis sativa), and Sunn hemp
(Crotalaria juncea) are potent source of fibers for textile
industries. This communication reports on the purifica-
tion and characterization of pectin lyase secreted by P.
citrinum MTCC 8897 and retting of L. usitatissimum, C.
sativa, and C. juncea fibers by the purified PNL.
ISSN 0006-2979, Biochemistry (Moscow), 2009, Vol. 74, No. 7, pp. 800-806. © Pleiades Publishing, Ltd., 2009.
Published in Russian in Biokhimiya, 2009, Vol. 74, No. 7, pp. 985-992.
800
Abbreviations: PNL, pectin lyase; SSF, solid-state fermentation.
* To whom correspondence should be addressed.
Purification and Characterization of Pectin Lyase
Secreted by Penicillium citrinum
S. Yadav
1
*, P. K. Yadav
2
, D. Yadav
2,3
, and K. D. S. Yadav
1
1
Department of Chemistry, D. D. U. Gorakhpur University, Gorakhpur 273009, India; E-mail: sangeeta_rahul@rediffmail.com
2
Department of Biotechnology, D. D. U. Gorakhpur University, Gorakhpur 273009, India
3
Department of Molecular Biology and Genetic Engineering, College of Basic Sciences and Humanities,
G. B. Pant University of Agriculture and Technology, Pantnagar (Uttarakhand) 263145, India
Received August 5, 2008
Revision received October 20, 2008
Abstract—The importance of various parameters such as sugarcane juice concentration, pH of the medium, and effects of
different solid supports for maximum secretion of pectin lyase from Penicillium citrinum MTCC 8897 has been studied. The
enzyme was purified to homogeneity by Sephadex G-100 and DEAE-cellulose chromatography. The molecular mass deter-
mined by SDS-PAGE was 31 kDa. The K
m
and k
cat
values were found to be 1 mg/ml and 76 sec
–1
, respectively. The opti-
mum pH of the purified pectin lyase was 9.0, though it retains activity in the pH 9.0-12.0 range when exposed for 24 h. The
optimum temperature was 50°C, and the pectin lyase was found to be completely stable up to 40°C when exposed for 1 h.
The purified pectin lyase was found efficient in retting of Linum usitatissimum, Cannabis sativa, and Crotalaria juncea.
DOI: 10.1134/S0006297909070141
Key words: pectin lyase, retting, solid state fermentation, Penicillium citrinum, Cannabis sativa, Crotalaria juncea, Linum usi-
tatissimum