1. Introduction
The Middle Triassic formed a crucial time for the
evolution of tetrapods, notably the archosaurs which
rose to the dominating land vertebrates during the
Early Triassic. However, unlike aquatic and amphi-
bious tetrapods, such as temnospondyls and phyto-
saurs, most archosaurs were terrestrial and thus not
often preserved with articulated skeletons. In the last
decade, focused research has yielded a range of new
archosaur and archosauriform taxa from Triassic de-
posits around the world, with Arizonasaurus, Effigia,
Dromomeron, Postosuchus, and Vancleavea from the
United States (NESBITT 2005, 2007; IRMIS et al. 2007;
PEYER et al. 2008; NESBITT et al. 2009a, b), Batracho-
tomus from Germany (GOWER 1999), Osmolskina,
Polonosuchus, Silesaurus, and an unnamed theropod
from Poland (DZIK 2003; SULEJ 2005; DZIK et al.
2008; BRUSATTE et al. 2009; BORSUK-BIA/ LYNICKA &
EVANS 2009), Hypselorhachis and Asilisaurus from
Tanzania (BUTLER et al. 2009; NESBITT et al. 2010),
and Qianosuchus from the southern Chinese Pro-
vinces Guizhou and Yunnan (LI et al. 2006).
In the last decade, excavations in the Middle
Triassic of southwestern Germany have yielded much
new material from the deposits of the Lower Keuper
or Lettenkeuper (Erfurt Formation). This formation
extends across most of the south and centre of
Germany, and has been long known for its richness in
vertebrate finds, although the mass abundance of the
large amphibian remains had long overshadowed the
actual diversity of other tetrapods (J AEGER 1824,
1828; MEYER & PLIENINGER 1844; SCHMIDT 1928;
WILD 1980). Whereas these marine-influenced swamp
deposits are extremely rich in temnospondyls and
other aquatic vertebrates (HELLRUNG 2003; SCHOCH
2006, 2008; WITZMANN et al. 2008), terrestrial tetra-
pods remained exceptional finds until very recently.
New archosauriform remains from the German Lower Keuper
Rainer R. Schoch
With 8 figures
SCHOCH, R.R. (2011): New archosauriform remains from the German Lower Keuper. – N. Jb. Geol.
Paläont. Abh., 260: 87–100; Stuttgart.
Abstract: Isolated bones of archosauriforms form regular finds in deposits of the Lower Keuper
(Erfurt Formation: upper Middle Triassic) of southwestern Germany. In addition to the fairly well
known 3-5 m long rauisuchian Batrachotomus kupferzellensis and the very fragmentary Zanclodon
laevis, remains of further, 1-2 m long archosauriforms have been identified. Although the number
of taxa represented by this incomplete material is unknown, the distinctness of these finds from
Batrachotomus and Zanclodon is indicated by the following features: (1) a “rauisuchian” type
maxilla with well-developed interdental plates, which forms a long suture with a posterodorsal
process of the premaxilla, (2) relatively long cervical and anterior dorsal vertebrae, (3) a gracile
humerus with a narrow proximal head and a weak deltopectoral crest, (4) a slender and markedly
curved tibia, and (5) tear-drop shaped, keeled armour plates. These findings highlight the diversity of
large predators in Middle Triassic land vertebrate faunas of Central Europe.
Key words: Archosauria, Archosauriformes, Erfurt Formation, Kupferzell, Ladinian, Vellberg.
©2011 Schweizerbart’sche Verlagsbuchhandlung, Stuttgart, Germany www.schweizerbart.de
DOI: 10.1127/0077-7749/2011/0133 0077-7749/2011/0133 $ 4.50
N. Jb. Geol. Paläont. Abh. 260/1, 87–100 Article
published online March 2011