Novel multifunctional acyloxyalkyl ester prodrugs of 5-aminolevulinic
acid display improved anticancer activity dependent on
photoactivation
G. Berkovitch
a
, A. Nudelman
b
, B. Ehenberg
c
, A. Rephaeli
d
, and Z. Malik
a
,
a
The Mina and Everard Goodman Life Sciences Faculty,
b
Chemistry Department,
c
Physics
Department, Bar Ilan University, Ramat Gan Israel;
d
Laboratory of Pharmacology and Experimental
Oncology, Felsenstein Medical Research Center, Sackler Faculty of Medicine, Tel-Aviv University,
ISREL
ABSTRACT
New approaches to PDT using multifunctional 5-aminolevulinic acid (ALA) based prodrugs activating mutual routes of
toxicity are described. We investigated the mutual anti-cancer activity of ALA prodrugs which upon metabolic
hydrolysis by unspecific esterases release ALA, formaldehyde or acetaldehye and the histone deacetylase inhibitor
(HDACI) butyric acid. The most potent prodrug in this study was butyryloxyethyl 5-amino-4-oxopentanoate (AN-233)
that stimulated a rapid biosynthesis of protoporphyrin IX (PpIX) in human glioblastoma U-251 cells and generated an
efficient photodynamic destruction. AN-233 induced a considerable high level of intracellular ROS in the cells following
light irradiation, reduction of mitochondrial activity, dissipation of the mitochondrial membrane potential resulting in
necrotic and apoptotic cell death. The main advantage of AN-233 over ALA stems from its ability to induce photo-
damage at a significantly lower dose than ALA.
Keywords: ALA-prodrugs, PpIX, PDT, glioblastoma U-251
1. INTRODUCTION
Glioblastoma (GBM) is the most malignant among astrocytic tumors and is associated with a poor prognosis.
Improvements in overall survival of patients with malignant gliomas have not been achieved in spite of intensive
treatment, including surgical resection, postoperative radiation therapy, and chemotherapy. Photodynamic therapy (PDT)
is a promising type of local therapy, and favorable results have been reported in several studies of PDT in glioma
patients
1-4
.
The use of endogenous PpIX generated though the heme biosynthetic pathway after administration of (ALA) has led to
many applications of PDT in cancer
5
. PpIX is a potent photosensitizer that accumulates excessively when ALA is given
externally to cells. The enzymes porphobilinogen deaminase (PBGD) and ferrochelatase (FC) are rate limiting and
contribute to the differential higher accumulation of PpIX in tumor cells
5, 6
. Subsequent activation of PpIX by light leads
to the formation of highly reactive singlet oxygen that causes the destruction of target cells by a complex cascade of
chemical, biological, and physiological reactions. A main disadvantage ALA-PDT is the hydrophilic nature of ALA
which limits its ability to penetrate though the skin or cell membranes. One solution to the poor penetrability of ALA is
to increase its bioavailability via the formation of esters
7
.
Our recent investigations revealed the anticancer activity of new acyloxyalkyl ester prodrugs of ALA
8
. The examination
of these prodrugs was evaluated mainly under dark conditions and showed that ALA prodrugs that release formaldehyde
(AN-243 and AN-242) were substantially more effective than the ALA- Acetaldehyde prodrug AN-233 in inducing
mortality of human glioblastoma, U-251 cells. These results were consistent with previous studies in which the
formaldehyde was shown to be a critical antiproliferative factor
9
. However, initial investigation of these prodrugs under
photo-irradiation conditions showed that both AN-233 and AN-243 that release ALA and butyric acid substantially
improved induction of cell mortality compared to ALA. AN-242 that releases ALA, formaldehyde and pivalic acid had
no improved activity with PDT. The enhanced effect of PDT mediated by both AN-233 and AN-243 compared to ALA
alone suggests that butyric acid, which is a component of both prodrugs, plays a role in improving the activity, while the
formaldehyde had no contribution.
Photodynamic Therapy: Back to the Future, edited by David H. Kessel, Proc. of SPIE Vol. 7380,
73800Z · © 2009 SPIE · CCC code: 1605-7422/09/$18 · doi: 10.1117/12.822703
Proc. of SPIE Vol. 7380 73800Z-1