Furanones in Strawberries: Evolution during Ripening and
Postharvest Shelf Life
Ana G. Pe ´rez, Raquel Olı ´as, Carlos Sanz, and Jose ´ M. Olı ´as*
Departamento de Fisiologı ´a y Tecnologı ´a de Productos Vegetales, Instituto de la Grasa, CSIC,
Avenida Padre Garcı ´a Tejero 4, 41012 Sevilla, Spain
Furaneol, mesifurane, and furaneol glucoside contents during ripening of four strawberry varieties
(Oso Grande, Chandler, Tudla, and I-101) have been analyzed. Patterns of furanone contents were
similar for the four varieties, amounts increasing during ripening to reach the highest values at
the overripe stage. However, furaneol and derivatives differed quantitatively among varieties. The
amount of furaneol shown by Oso Grande at the overripe stage is the highest so far reported (37.05
μg/g fw). The highest content in mesifurane and furaneol glucoside was found in the I-101 variety,
at the overripe stage, 23.5 and 13.2 μg/g fw, respectively. Results obtained in eight different varieties,
at commercial maturity stage, also showed quantitative differences. Strawberries were stored at 1
°C for 2 days to simulate refrigerated transport and then kept for 7 days at 17 °C to simulate the
shelf life period. At 17 °C, the amount of mesifurane and furaneol glucoside increased more than
50% with concomitant loss of furaneol.
Keywords: Strawberry; flavor; furanones
INTRODUCTION
Aroma development is one of the most prominent
changes that occurs during ripening of fruit. The aroma
is also an important quality factor that influences
consumer acceptability of fruits. Strawberry aroma is
mainly determined by a complex mixture of esters,
alcohols, aldehydes, and sulfur compounds (Schreier,
1980; Dirinck et al., 1981). Although no character
impact compound has been found for strawberry aroma,
some researchers consider 2,5-dimethyl-4-hydroxy-3(2H)-
furanone (furaneol) and 2,5-dimethyl-4-methoxy-3(2H)-
furanone (mesifurane) as two of the most important
aroma contributors (Pyysalo et al., 1979; Larsen and
Poll, 1990, 1992). Both furaneol and mesifurane have
strong, sweet, and pleasant odors. Furaneol imparts
caramel burnt sugar notes at high concentrations and
becomes fruity at lower concentrations (Re et al., 1973).
Mesifurane is described as having a more sherry-like
aroma (Hunter et al., 1974). Recently Larsen and Poll
(1992) found that a mixture of furaneol and ethyl
butanoate presented a strawberry-like odor.
Several studies have identified the presence of fura-
neol, mesifurane (Hirvi and Honkanen, 1982; Douillard
and Guichard, 1990), and furaneol glucoside (Mayerl et
al., 1989) in strawberries, but these compounds have
not been found in all cultivars. Factors such as furaneol
water-soluble nature (Pyysalo et al., 1979; Douillard and
Guichard, 1989) and thermal instability (Flath and
Forry, 1970; Shu et al., 1985) could well account for the
failure of some authors to detect these compounds.
The new analysis procedure described by Sanz et al.
(1994) involves HPLC separation and quantitation of
furaneol and its derivatives which avoids some of the
risks of thermal oxidative decomposition inherent in GC
analysis. In this work, changes in furanone contents
during strawberry ripening and postharvest shelf life
have been studied using this method.
EXPERIMENTAL PROCEDURES
Materials. Plants from Oso Grande, Chandler, Tudla, and
I-101 strawberry varieties were greenhouse grown in optimum
conditions of light, soil, and diseases control. Strawberry
flowers were marked the day of blooming, and fruits were
harvested at four visual ripening stages: white, pink, red, and
dark-red (overripe). Strawberry fruits from Camarosa, Car-
tuno, Cartcua, Carlsbad, Laguna, Sunset, Seascape, and
Cuesta varieties were grown in field at Cartayfres (Cartaya,
Huelva) and sampled at commercial maturity stage (more than
75% red color). Oso Grande fruits used for the shelf life study
were also harvested at commercial maturity stage in Torreagro
(San Bartolome de las Torres, Huelva). Fruits were placed in
0.5 kg poly(ethylene) terphthalate baskets mechanically filmed
with poly(propylene) (Poligal, thickness 25 μm), by means of
a flow-pack device. The poly(propylene) film had an O
2
transmission rate of 750 cm
3
/m
2
/24 h at 23 °C and a moisture
vapor trasmission rate of 1.1 g/m
2
/24 h at 23 °C and 85% RH.
Fruits were immediately stored at 1 °C for 2 days to simulate
refrigerated transport and then stored for 7 days at 17 °C to
simulate the shelf life period. Control fruits were stored at 1
°C.
Preparation of Sample for HPLC. Strawberries were
cut symmetrically in four pieces. Four portions from four
different fruits (ca. 20 g) were randomly sampled and ground
with 5 mL of distilled water in a Waring blender at 0-4 °C.
Celite 545 (10 g) was added and after mixing allowed to settle
for 5 min. The mixture was filtered, washed three times with
10 mL of distilled water, and again filtered first through a
0.45 μm and then through a 0.2 μm nylon membrane before
HPLC analysis.
HPLC Analysis. Quantitative HPLC analysis was carried
out according to the method described by Sanz et al. (1994)
with slight modifications. A liquid chromatograph, Beckman
Golden System, equipped with an ODS (4.6 mm × 250 mm) 5
μm column was used. UV detector was selected at 280 nm,
and the injection volume was 20 μL. The mobile phase utilized
for the separation of furanones consisted of two eluents: 0.2
M sodium acetate/acetic acid, pH 4 (solvent A), and methanol
(solvent B). Chromatographic conditions were 0-11 min,
isocratic 13% B; 11-26 min, gradient 13-23% B; 26-30 min,
isocratic 23% B; and 30-33 min, gradient 23-80% B (cleaning
process).
* Author to whom correspondence should be ad-
dressed.
3620 J. Agric. Food Chem. 1996, 44, 3620-3624
S0021-8561(96)00099-4 CCC: $12.00 © 1996 American Chemical Society