European Journal of Epidemiology 10:707-711, 1994.
© 1994 Kluwer Academic Publishers. Printed in the Netherlands.
Serotypes of CNFl-producing Escherichia coli strains that cause
extraintestinal infections in humans
J. E. Blanco 1, J. Blanco 1, M. Blanco 1, M. P. Alonso 2 & W. H. Jansen 3
t Departamento de Microbioloxia e Parasitoloxia, Facultade de Veterinaria, Universidade de Santiago, Lugo, Spain;
2 Unidade de Microbioloxia, Complexo Hospitalario Xeral-Calde, Lugo, Spain; 3Laboratory of Bacteriology and
Antimicrobial Agents, National Institute of Public Health and Environmental Protection, Bilthoven, The Netherlands
Accepted in revised form 15 July 1994
Abstract. The O:K:H serotypes of 137 neerotoxi-
genic Escherichia coli (NTEC) producing the cyto-
toxic necrotizing factor type 1 (CNF1) isolated from
human extraintestinal infection were determined.
Although NTEC producing CNF1 belonged to 58
different serotypes, only 10 of them accounted for
54% of strains. The most common serotypes, in order
of frequency, were: O4:K?:H5, O6:KI3:H1,
O83:Kl:H31, O75:K95:H5, O2:Kl:H6, O2:K7:H-,
O75:Kl:H7, O2:K?:H1, O4:K12:H1 and
O22:K13:H1. CNF1 strains of serotypes O2:K7:H-
and O4:K12:H1 express P-fimbriae, whereas CNF1
strains of serotypes O2:K?:H1, O2:Kl:H6 and
O75:K95:H5 possess the adhesin responsible for
MRHA type III. Among CNF1 strains of serotype
O4:K?:H5 there exist some that express P-fimbriae
and others that possess MRHA type III. Lastly, the
majority of CNF1 strains of serotypes O6:K13:H1,
O22:KI3:H1, O75:Kl:H7 and O83:Kl:H31 do not
express P-fimbriae nor the adhesin responsible to
MRHA type III. Our results show that extraintestinal
infections are caused by a limited number of virulent
clones, as suggested by the theory of special patho-
genicity.
Key words: Necrotoxigenic Escherichia coli, Pathogenicity, Toxins, Virulence
Introduction
Necrotoxigenic Escherichia coli (NTEC) strains are
able to elaborate two types of cytotoxic necrotizing
factors (CNF1 and CNF2) that induce cell multinu-
cleation in Vero and HeLa tissue cultures, cause
necrosis in rabbit skin and are lethal to mice [1 ]. Both
necrotic toxins are cell-associated products easy to
detect when bacteria are sonicated or when grown
in the presence of mitomycin C [2]. CNF1 and CNF2
are heat-labile proteins of 115 and 110 kilodaltons,
respectively [3-6]. Whereas CNF1 is encoded by
chromosomic genes [7], CNF2 is encoded by trans-
missible plasmids [5]. NTEC were associated with
human extraintestinal infections [8-13] and isolated
from calves with diarrhoea or septicaemia [14-16].
NTEC of human origin usually produce CNF1,
whereas bovine NTEC generally synthesize CNF2
[17]. We have recently established that the majority
of human CNF1 strains belong to serogroups 02, 04,
06, O14, 022, 075 or 083, and that the bovine
NTEC producing CNF2 usually belong to serogroups
O1, 03, O15, 055, 088 or O123 [17]. The present
report is the first one to describe the O:K:H serotypes
of CNF1 E.coli strains that cause human extrain-
testinal infections in Spain.
Materials and methods
Bacterial strains. A total of 137 CNF1 NTEC strains
isolated from patients with extraintestinal infections
between 1986 and 1992 were investigated, Strains
from asymptomatic bacteriuria, cystitis, pyelo-
nephritis, bacteremia, peritonitis, appendicitis, chole-
cystitis, wound and respiratory infections and other
miscellaneous sources were isolated from both out-
patients and inpatients attending three hospitals in
Galicia (northwestern Spain): Residencia Sanitaria
Juan Canalejo of La Corufia, Hospital Xeral-Calde of
Lugo and Hospital Comarcal da Costa of Burela. A
single E. coli strain was obtained from each patient.
Isolation and identification of E. coli was performed
by standard bacteriological methods. Reference E.
coli strains used as positive and negative controls
were: MR48 (O75:K95, MRHA III ÷, CNF1 ÷, Hly+),
MR199 (O6:K13, MRHA HI+, CNF1 ÷, Hly+), C1212-
77 (O6:K2:H1, P-fimbriated) and C1976-79
(OI:KI:H7, P-fimbriated). Strains were stored at
room temperature in nutrient broth (Difco, USA) with
0.75% of agar.
Production and detection of CNF1. For the detec-
tion of CNF1 the filtrates of cultures treated with
mitomycin C were assayed on Vero and HeLa cells
as previously described [2]. The Vero and HeLa cell