160 Journal of Basic Microbiology 2008, 48, 160 – 167
© 2008 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim www.jbm-journal.com
Research Paper
Production of an extracellular thermohalophilic lipase from
a moderately halophilic bacterium, Salinivibrio sp. strain SA-2
Mohammad Ali Amoozegar
1
, Ensieh Salehghamari
1
, Khosro Khajeh
2
, Mahbube Kabiri
1
and Saied Naddaf
3
1
Extremophiles Lab., Dept. of Microbiology, Faculty of Biology, College of Science, University of Tehran,
Tehran, Iran
2
Department of Biochemistry, Faculty of Science, Tarbiat Modarres University, Tehran, Iran
3
Parasitology department, Pasteur Institute of Iran, Teheran, Iran
Fifty strains of moderately halophilic bacteria were isolated from various salty environments in
Iran. A strain designated as SA-2 was shown to be the best producer of extracellular lipase and
was selected for further studies. Biochemical and physiological characterization along with 16S
rDNA sequence analysis placed SA-2 in the genus Salinivibrio. The optimum salt, pH,
temperature and aeration for enzyme production were 0.1 M KCl, pH 8, 35 °C and 150 rpm,
respectively. The enzyme production was synchronized bacterial growth and reached a
maximum level during the early-stationary phase in the basal medium containing 1 M NaCl.
Triacylglycerols enhanced lipase production, while carbohydrates had inhibitory effects on it.
The maximum lipase activity was obtained at pH 7.5, 50 °C and CaCl
2
concentration of 0.01 M.
The enzyme was stable at pH range of 7.5 – 8 and retained 90% of its activity at 80 °C for
30 min. Different concentrations of NaNO
3
, Na
2
SO
4
, KCl and NaCl had no affect on lipase
stability for 3 h. These results suggest that the lipase secreted by Salinivibrio sp. strain SA-2 is
industrially important from the perspective of its tolerance to a broad temperature range, its
moderate thermoactiviy and its high tolerance to a wide range of salt concentrations (0 – 3 M
NaCl).
Keywords: Halophilic bacteria / Lipase activity / Lipase production / Salinivibrio / Thermohalophilic enzyme
Received: November 05, 2007; accepted: January 09, 2008
DOI 10.1002/jobm.200700361
Introduction
*
Lipases are among the most important hydrolytic en-
zymes with potential applications in a variety of bio-
technological fields such as food, dairy, detergent
pharmaceutical, agrochemical and oleochemical indus-
tries. Many microbial lipases are available now and
some of them have already been used in various indus-
tries. However, they are not sufficient to meet most of
the industrial demands. Since industrial processes are
commonly carried out under harsh conditions, it would
be of great importance to produce types of enzymes
Correspondence: M. A. Amoozegar, Extremophiles Lab., Dept. of Micro-
biology, Faculty of Biology, College of Science, University of Tehran,
P.O. Box 14155-6455, Tehran, Iran
E-mail: amozegar@khayam.ut.ac.ir
Phone: +98-21-61112622
Fax: +98-21-66492992
which retain their optimal activity at extremes of pH,
temperature and different concentrations of salts.
Moderately halophilic bacteria are extremophiles mi-
croorganisms which have adapted to live in a wide
range of NaCl (1 – 25%) and constitute an interesting
group of microorganisms that could be used as a source
of such enzymes [1]. These organisms produce halotol-
erant enzymes which are able to tolerate high ionic
strength (up to 4 M NaCl) in their environments and in
contrast to their extremely halophilic counterparts can
be active in the absence of salts. Recently, a consider-
able attention has been given to the enzymes produced
by moderately halophilic microorganisms and their
biotechnological potentials [2]. While there are numer-
ous reports on the lipase production and characteriza-
tion from non-halophilic microorganisms, very limited
information is available on lipases from halophilic spe-