Immunopharmacology, 7 (1984) %16
Elsevier 9
IMO 00211
Immunochemical Studies on Thymosin: Radioimmunoassay
of Thymosin f14
Paul H. Naylor*, John E. McClure**, Bryan L. Spangelo, Teresa L.K. Low and Allan L. Goldstein
Department of Biochemistry, The George Washington University School of Medicine, 2300 Eye Street, N.W., Washington, D.C. 20037,
U.S.A.
(Received 2 June 1983;accepted after revision 15 August 1983)
Abstract: Thymosinf14,a peptide with hormonal-like properties first isolated from the thymus gland, can be measured in serum using
a newly described radioimmunoassay. The radioimmunoassay utilizes an antibody raised in rabbits against synthetic thymosin t4
conjugated by glutaraldehyde to keyhole limpet hemocyanin. A l z5i_tyrosine.Cl3 analogue of the biologicallyactive C-terminal frag-
ment is used as the radioactive tracer. The radioimmunoassayis sensitive in the nanogram range and no cross-reactivitywith common
serum proteins is demonstrable. High performance liquid chromatography of serum samples indicates that two thymosin t , cross-
reactive species are present in human serum. Levelsin serum range from 450 to 1100ng/ml and decline with age.
Key words: Thymosinf14;Thymichormone; Radioimmunoassay
Introduction
One of the most important advances in medical re-
search during the past decade has been the devel-
opment of our understanding of the role played by
the thymic-dependent immune system in various
diseases including the primary immunodeficiency
diseases, cancer, infectious diseases, and the au-
toimmune diseases (White et al., 1975; Goldstein et
al., 1970; Goldstein et al., 1981; Low et al., 1982).
Our research program has focused upon studies
designed to elucidate the endocrine role of the thy-
mus gland and has resulted in the identification of
a family of polypeptides from the thymus which
have been collectively termed thymosins (Goldstein
et al., 1981; Low et al., 1982).
Thymosin t4 is a 4963 molecular weight poly-
peptide that we have isolated from calf thymus
(Low et al., 1981). The amino acid sequence of thy-
mosin 1/4 has been determined and published (Low
et al., 1982); this polypeptide induces the expression
of terminal deoxynucleotidyl transferase in trans-
ferase-negative murine lymphocytes in vivo and in
vitro (Low et al., 1981). Thymosin 1/4 has also been
found to elevate luteinizing hormone releasing hor-
mone (LHRH) and luteinizing hormone (LH) in an
* To whom reprint requests should be addressed.
** Present address: Section of Allergyand Immunology,Baylor College of Medicine, Houston, TX 77030, U.S.A.
Abbreviations: LHRH: luteinizing hormone releasing hormone; LH: luteinizing hormone; RIA: radioimmunoassay; thymosin F5:
thymosin fraction five; RIAB: radioimmunoassaybuffer; IgG: immunoglobulinG; cpm: counts per minute; TFA: trifluoroacetic acid;
HPLC: high performance liquid chromatography.
0162-3109/84/$03.00 © 1984 Elsevier Science Publishers B.V.