Tissue & Cell 1998 30 (5) 525-530
© 1998 Harcourt Brace & Co. Ltd
Effect of culture conditions on endo-
thelial cell growth and responsiveness
Ingrid A. M. Relou', Cora A. Damen 1, Daisy W. J. van der Schaft 2,
Gerard Groenewegen, Arjan W. Griffioen 2
Abstract. The in vitro culture of endothelial cells (EC) is dependent on the presence of a coated surface and the
availability of growth factors in the medium. The aim of the present research is to investigate whether in vitro EC
culture conditions, such as serum source and surface coating, determine the growth characteristics of EC. The
phenotype of EC was studied at the level of adhesion molecule expression and down-regulation by angiogenic
factors. We found that human umbilical vein EC adhere well to and stretch well with plastic coated with
fibronectin, collagen, gelatin and hyaluronan in contrast to non-coated plastic. While low in hyaluronan-coated
wells, the spontaneous proliferation of EC was enhanced in fibronectin-collagen and gelatin-coated wells as
compared to non-coated wells. Basic fibroblast growth factor bFGF-induced proliferation, however, was best on
hyaluronan-coated plastic. A markedly up-regulated proliferation was measured on fibronectin and collagen while
EC on gelatin-coated plastic only showed moderate bFGF-induced proliferation. On non-coated plastic EC were
not inducible with bFGF. The induction of apoptosis by serum deprivation on these different matrices was most
efficient when no coat was available or when wells were coated with hyaluronan, and bFGF inhibited apoptosJs
induction under all conditions. The use of different culture media demonstrated that human and bovine serum
both can be used for human EC assays. The synthetic medium Utroser G prevented both spontaneous and
growth factor-induced proliferation. We found that apart from some magnitude differences, the down-regulation of
intercellular adhesion molecule-1 (ICAM-1) by angiogenic factors such as bFGF is not dependent on specific
culture conditions.
Keywords: Endothelial cell, extracellular matrix, adhesion molecules, angiogenesis, in vitro
Introduction
The extracellular matrix of endothelial cells (EC) is a dense
network composed of macromolecules such as collagen,
fibronectin, vitronectin, hyaluronan, laminin and glycosaln-
inoglycans. The interaction of EC with these components by
specific receptors plays an important role in the growth and
~Department of Internal Medicine and Medical Oncology, Laboratory for
Angiogenesis Research, University Hospital Utrecht, Heidelberglaan 100, PO
Box 85500, 3508 GA Utrecht, The Netherlands. 2Departmentof Internal
Medicine, Tumor Angiogenesis Laboratory University Hospital Maastricht,
PO Box 5800, 6202 AZ Maastricht, The Netherlands.
Received 6 January 1998
Accepted 9 April 1998
Correspondence to: Dr A.W. Griffioen. Fax: +31 43 3875006; E-mail:
a.griffioen @intmed.unimaas.nl
differentiation of EC and vascular development, e.g. during
tumor progression (Liotta, 1986; Dejana et al., 1987; Ingber,
1993; Ingber et al., 1995).
EC growth and differentiation is also determined by
specific growth factors. Outgrowth of tumors is dependent
on the formation of new vasculature and therefore malignant
cells produce high levels of angiogenic factors (Folkman,
1990; Folkman & Shing, 1992). We found that these angio-
genesis factors down regulate endothelial adhesion molecule
expression thereby preventing extravasation and infiltration
of leukocytes (Griffioen et al., 1996a, 1996b). To investigate
the role of the extracellular matrix in these phenomena we
studied the spontaneous and growth factor-induced prolifera-
tion of EC in vitro on different types of matrix protein-coated
plastic and in the presence of different culture media. In
addition, the induction of apoptosis by serum deprivation
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