Gen. Pharmac. Vol. 27, No. 8, pp. 1373-1376, 1996
Copyright © 1996 Elsevier Science Inc.
Printed in the USA.
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ISSN 0306-3623/96 $15.00 + .00
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Selective Effect of
2-(Polyhydroxyalkyl)-Thiazolidine-4-Carboxylic Acids
on Nonprotein Sulfhydryl Groups in Tumor Bearing Mice
L. Wtodek,* M. Wr6bel, and J. Czubak
INSTITUTE OF MEDICALBIOCHEMISTRY, COLLEGIUMMEDICUM,
JAGIELLONIAN UNIWRSITY, 7 KOI'ERNI~ STREET, 31-034 KRAKdw, POLAND [FAx: (48-12) 21-46-64]
ABSTRACT. 1. Thiazolidine derivatives (TD), the products of condensation of L-cysteine (cys)
with sugars (D-glucose, D-xylose, D-arabinose, D-galactose, and D-mannose), successfully elevated non-
protein sulfhydryl (NPSH) levels in livers of Ehrlich ascites tumor (EAT)-bearing mice.
2. At the same time, TD promoted a significant drop of NPSH in EAT cells.
3. Thus, TD, through their selective influence on the levels of NPSH in liver and cancer cells ap-
pear to be promising compounds for anticancer therapy. Copyright © 1996 Elsevier Science Inc. GEN
eHARMAC 27;8:1373--1376, 1996.
KEY WORDS. L-Cysteine, nonprotein sulfhydryl groups, thiazolidine derivatives, Ehrlich ascites tumor
INTRODUCTION
Glutathione (GSH) constitutes an almost complete pool of nonpro-
tein sulfhydryl compounds (NPSH) in the cells; it plays an impor-
tant role as a cellular protector (Meister, 1989; Reed, 1990). GSH
is known as a substrate for glutathione S-transferase (EC 2.5.1.18)
and glutathione peroxidase (EC 1.11.1.9), enzymes catalyzing reac-
tions of detoxification of xenobiotic compounds and antioxidation
of reactive species and free radicals (Bray and Taylor, 1993; Meister,
1991). Cellular GSH alters the toxicity of many chemotherapy
drugs and radiation (Bray and Taylor, 1993; Durand and Olive,
1989; Meister, 1991). It has been suggested that a modulation of
GSH levels would be a promising therapeutic strategy (Anderson
and Meister, 1987; Meister, 1985; Meister, 1991). This is the reason
for the intensive search for such compounds which, through the in-
crease of GSH levels in normal cells, protect them but at the same
time offer no protection for tumor cells. Obviously, the ideal drugs
would be those compounds that could simultaneously elevate the
GSH level in normal cells and decrease it in tumor cells.
GSH alone cannot be used as a drug because it does not enter the
cell (Hahn et al., 1978). Free L-cysteine (cys), as an amino acid lim-
iting the biosynthesis of GSH, is very toxic to the central nervous
system (Karlsen et al., 1981; Olney et al., 1971). Methionine, being
a source of cys through the cystathionine pathway, decreases the
level of ATP in the cells (Hardwick et al., 1970), and is not a good
source of cys for GSH synthesis in the liver (Stipanuk et al., 1992).
As shown in our previous studies, thiazolidine derivatives (TD) are
able to increase the concentration of NPSH in the livers of healthy
mice (Wtodek et al., 1993). These compounds can gradually release
cys by enzymatically controlled processes or by nonenzymatic hydro-
lysis (Cavallini et al., 1956; Nagasawa et al., 1984, Roberts and
Francetic, 1991; Wtodek et al., 1993) (Scheme 1).
Furthermore, as reported by Roberts and Francetic (1991), the
condensation product of t-cysteine with D-ribose selectively ele-
vated GSH levels in numerous organs of tumor-bearing mice. This
is why we investigated the effect of several 2-(polyhydroxyalkyl)-
thiazolidine-4-carboxylicacids, condensation products of i.-cysteine
*To whom correspondence should be addressed.
Received 26 October 1995; accepted 9 January 1996.
with sugars, on the level of NPSH and cys in EAT cells and in the
livers of tumor-bearing mice.
MATERIALS AND METHODS
Chemicals
The following thiazolidine derivatives (TD), the condensation products
of c-cysteine with sugars (D-glucose, D-xylose, D-arabinose, D-galactose,
D-mannose), were used in our experiments.
1. 2-D-glucopentahydroxypentyl)-thiazolidine-4-carboxyiic acid (CGlc).
2. 2-(D-xylotetrahydroxybutyl)-thiazolidine-4-carboxylic acid (CXyl).
3. 2-(D-arabinotetmhydroxybutyl)-thiazolidine-4-carboxylic acid (CAm).
4. 2-(D-galactopentahydroxypentyl)-thiazolidine-4-carboxylic acid (CGal).
5. 2-(D-mamaopentahydroxypentyl)-thiazolidine-4-carboxylic acid (CMan).
Compound numbers 3, 4, and 5 were synthesized by method A,
described by Radomski and Temeriusz (1989). Compound numbers
1 and 2 were synthesized by a modified method of Weitzel et al.
(1959) in the Laboratory of Organic Synthesis, Jagiellonian Univer-
sity (Krak6w, Poland).
c-Cysteine, reduced glutathione, 5,5-dithiobis-(2-nitrobenzoic
acid) (DTNB), and bovine serum albumin were provided by Sigma
Chemical Co., and ninhydrin was from Fluka AG.
The EATC were maintained in Swiss female mice, 3 months of
age, with an approximate weight of 20 g, kept on a standard mouse
diet. Seven days after implantation of EATC, the animals were
given intraperitoneal injections of thiazolidine derivatives at the
dose of 1.2 minol/kg, once a day for 3 days. The control animals were
injected with the same volume of isotonic saline. At day 4, the mice
were killed and their livers were dissected. EATC were collected
and washed by repeat suspensions in saline. Subsequently, the livers
and EATC were frozen at -30°C until needed. After thawing, tu-
mor cells and livers were homogenized with 0.1 M phosphate buffer,
pH 7.4 (1 g:4 mL), and centrifuged at 650 g for 15 rain to determine
the levels of NPSH and cys in the supernatant fractions.
NPSH was measured by reaction with 5,5-dithiobis-(2-nitroben-
zoic acid) (DTNB) to yield a compound that absorbs at 412 nm
(Moron et al., 1979). Cysteine was determined according to Gai-
tonde (1967). Protein content was measured according to the