Abstract: We investigated the impact of respiratory syncytial virus
(RSV) infection, an important asthma precipitant, on endothelin
receptor function and release in sheep bronchial explants. RSV
infection was confirmed using polymerase chain reaction and
immunohistochemistry. Since sheep airway smooth muscle contains
only endothelin-A receptors, sarafotoxin (Stx) S6c did not cause
airway contraction. In contrast, sarafotoxin S6c (300 nM) caused
contraction in RSV-infected bronchial explants (8 ± 3% carbachol
E
max
). However, we could not detect airway smooth muscle
endothelin-B receptors in explants using autoradiography. RSV
infection per se did not alter the release of immunoreactive
endothelin from sheep bronchial explants (control = 11.6 ± 0.9 pg
versus RSV = 12.1 ± 0.9 pg). Interestingly, dexamethasone (1 μM)
alone increased endothelin release in both control (17.9 ± 2.0 pg)
and RSV-infected tissue (18.3 ± 3.1 pg). The combined presence of
protease-activated receptor-2 (PAR-2) ligand (100 μM) and
dexamethasone (1 μM) also increased endothelin release from
control tissue (17.3 ± 1.4 pg), but endothelin release was suppressed
by PAR-2 ligand in RSV-infected tissue (10.3 ± 0.8 pg), probably
because PAR-2 expression was increased by RSV. In summary, the
novel expression of endothelin-B receptors triggered by RSV might
be relevant to RSV-associated asthma. Furthermore, activation of
airway PAR-2 may be protective in asthma where endothelin levels
are elevated in part via endothelin release suppression.
Key Words: respiratory syncytial virus, endothelin, bronchial
explants, airway smooth muscle
(J Cardiovasc Pharmacol™ 2004;44(suppl 1):S202–S206)
R
espiratory tract viral infections commonly precipitate
and exacerbate asthma symptoms. Indeed, 85% of
asthma exacerbations in children and about 50% of asthma-
like symptoms in adults involve respiratory tract viral
infection.
1
Most children have been infected with respiratory
syncytial virus (RSV) by the age of 2 years. The development
of bronchiolitis associated with RSV infection in infants
under the age of 1 years has been associated with an
increased risk for developing asthma.
2
There is a wealth of evidence to suggest that endothelin
(ET)-1 plays a mediator role in asthma. Importantly, in
asthma, immunoreactive ET levels are increased in airway
epithelium and in bronchoalveolar lavage fluid. In addition,
ET-1 is a potent bronchoconstrictor, a secretagogue for
bronchial mucous and an airway smooth muscle mitogen.
ET-1 also has pro-inflammatory actions in the airways,
stimulating the release of inflammatory cytokines.
3
We have previously shown that respiratory tract viral
infection with parainfluenza-1,
4
parainfluenza-3
5
and
influenza-A
6
is associated with changes in ET receptor
function and distribution in airway smooth muscle. We have
also shown that influenza-A virus infection with associated
airway inflammation markedly increased ET production in
mouse airways.
7
Importantly, RSV infection of airway
epithelial cells in culture significantly increased the
expression of ET-1 mRNA.
8
Lambs inoculated with a human strain of RSV develop
clinical signs of respiratory tract infection with an
immunological response to RSV present in the airways.
9
We
have developed an airway explant culture system using sheep
bronchi in order to establish infection with RSV.
10
The current
study examined the impact of RSV infection on ET receptor
function and distribution in sheep bronchial explants. In
addition, the release of immunoreactive ET from control and
RSV-infected bronchial explants was investigated.
METHODS
Explant Culture
Sheep (female and castrated male Merinos,
1–10 months old) were sacrificed by an experienced animal
care technician using a captive bolt and were exsanguinated
before the lungs were removed. Bronchial rings (3–6 mm,
internal diameter) were obtained and explants established.
10
Bronchial rings were placed individually in 35-mm plastic
culture dishes containing 2 mL serum-free CMRL 1066
S202 J Cardiovasc Pharmacol™ • Volume 44, Supplement 1, November 2004
SUPPLEMENT ARTICLE
The Impact of Respiratory Syncytial Virus Infection
on Endothelin Receptor Function and Release in
Sheep Bronchial Explants
Lynette B. Fernandes*, Angela C. D’Aprile*, Glenn J. Self*, Gerald B. Harnett†, and Roy G. Goldie*
*Pharmacology Unit, School of Medicine & Pharmacology, The University
of Western Australia and Western Australian Institute for Medical
Research, Perth, Western Australia, Australia, and †Department of
Microbiology and Infectious Diseases, The Western Australian Centre
for Pathology and Medical Research, Perth, Western Australia, Australia
Address correspondence and reprint requests to Dr Lynette Fernandes,
Pharmacology Unit (M510), School of Medicine and Pharmacology, The
University of Western Australia, 35 Stirling Highway, Crawley, WA
6009, Australia. E-mail: lfernandes@pharm.uwa.edu.au
This work was supported by the Asthma Foundation of Western Australia
Inc.
Copyright ©2004 by Lippincott Williams & Wilkins