1 23 Journal of Applied Phycology ISSN 0921-8971 Volume 28 Number 3 J Appl Phycol (2016) 28:1509-1520 DOI 10.1007/s10811-015-0726-y Impacts of nitrogen and phosphorus starvation on the physiology of Chlamydomonas reinhardtii Manoj Kamalanathan, Mattia Pierangelini, Lauren Ann Shearman, Roslyn Gleadow & John Beardall