NATURE BIOTECHNOLOGY VOLUME 24 NUMBER 4 APRIL 2006 411
The paths around stem cell intellectual property
Kenneth S Taymor, Christopher Thomas Scott & Henry T Greely
Can new approaches for deriving human stem cells circumnavigate existing patents that dominate embryonic stem cell
intellectual property?
I
n November 1998, James Thomson, of
the University of Wisconsin in Madison,
Wisconsin, reported the first derivation of
human embryonic stem (hES) cell lines
1
. Less
than two years earlier, Ian Wilmut (then at
Roslin Institute, Edinburgh, UK) had electrified
the world by announcing the birth of Dolly, the
first mammal cloned from an adult donor cell
2
.
Taken together the two discoveries promised
the possibility of making hES cell lines carefully
tailored to match a particular donor. Many
expected, and continue to expect, these discov-
eries to transform biomedical sciences.
But with the discoveries came patents.
Thomson’s work was the subject of two impor-
tant patents
3
, owned by the Wisconsin Alumni
Research Foundation (WARF); patents on
Wilmut’s work were assigned to Roslin Institute
and then to Geron (Menlo Park, CA, USA) when
that firm acquired Roslin
4
. Concerns have been
raised that the breadth of the patents’ claims
encompass all hES cell and downstream thera-
peutic and diagnostic products, regardless of the
stem cell derivation process employed
5
.
Both WARF—through its affiliate, WiCell
Research Foundation (WiCell)—and Geron
will permit other institutions or firms to use
these patents under individually negotiated
materials transfer agreements (MTAs)
6
. Those
MTAs, however, require the researchers to make
an initial cash payment and prohibit commer-
cial research without an additional license. The
commercial licenses require sharing profits with
WiCell or Geron. These ‘reach-through’ rights
are often cited as impediments to a free and clear
research environment, constraining the ability
of stem cell research to move forward
5
.
Since Thomson’s and Wilmut’s initial work,
researchers have explored refinements, varia-
tions and entirely new ways to produce stem
cell lines. Here, we explore these new methods
and the effect they have on understanding the
scope of the existing US patents. We first ana-
lyze the claims of Thomson’s patents on hES cell
lines and Wilmut’s somatic cell nuclear transfer
(SCNT) patents and then describe emerging
technologies for creating useful pluripotent cells.
Although we offer no definitive legal opinions,
we find that some new methods and the stem
cells they create may be outside the scope of the
Thomson and Wilmut patent claims. This has
important implications for understanding the
application, scope and constraints of the hES cell
patent landscape.
The core hES cell patents
The two key Thomson patents, issued in 1998
and 2001, will expire in the United States in 2015,
20 years from the earliest date of filing. Geron,
which funded Thomson’s work, has an exclu-
sive license to use the Thomson patents for three
important therapeutic and diagnostic areas: the
use of embryonic cell lines to make neural, car-
diac and insulin-producing cells. Any researcher
seeking to use the patents for research in those
areas would need a license from Geron.
The key Thomson patents claim both the
composition of stem cells and cell lines them-
selves as well as the process used to create them.
A key composition claimed in the Thomson pat-
ents is pluripotent hES cells with four significant
characteristics: the cells (i) proliferate in vitro
for over one year; (ii) maintain a stable, euploid
chromosome karyotype; (iii) have the poten-
tial to differentiate into the three germ layers,
endoderm, mesoderm and ectoderm; and (iv)
are inhibited from differentiation when cultured
on fibroblast feeder layers. Any ‘pluripotent hES
cells’ with these four characteristics fall within
that claim. Additional compositions claimed
include hES cells with the ability to differentiate
spontaneously to extra-embryonic tissues (the
trophoblast) and hES cells with certain specific
markers.
The Thomson Process claim describes creat-
ing hES cells by (i) growing a human embryo
to the blastocyst stage, (ii) removing cells of
the inner cell mass (ICM), (iii) plating them on
embryonic fibroblasts, (iv) dissociating the mass,
(v) replating the dissociated cells on embryonic
feeder cells, (vi) selecting colonies with compact
morphologies and cells with high nucleus-to-
cytoplasm ratios and prominent nucleoli, and
(vii) culturing the cells of the selected colonies
to obtain a stem cell line. Thomson claims any
cell lines created by this process regardless of
whether the cell line has the characteristics that
he described in his patents (See Figure 1).
The Wilmut patents on SCNT
Wilmut and his collaborators patented their
original work in nonhuman mammalian SCNT
in several countries, including the United States
4
.
The US patent, originally assigned to the Roslin
Institute, was transferred to Geron, and will
expire in 2016. Wilmut claims several variations
on a process for transferring a donor nucleus
into a recipient cell of the same species includ-
ing activation after nuclear transfer, incubation
of the SCNT product, and transferring and
developing the blastocyst in utero. The recipi-
ent cell may be an enucleated oocyte, zygote or
cell extracted after cleavage. The patent describes
several methods of enucleation, including physi-
cal (aspiration) and noninvasive (irradiation
with ultraviolet light).
The Wilmut patents address nonhuman uses
of SCNT for the production of animals, not
stem cells. Their relevance in hES cell research,
however, may be significant because the series
of techniques that they describe are one likely
means of producing hES cells by SCNT. In addi-
tion, commercialization of SCNT-derived hES
cells is likely to require use of SCNT animal
models and testing.
Kenneth Taymor, Christopher Thomas Scott
and Henry T Greely are at the Stanford
University Program on Stem Cells in Society,
Stanford University, 701 Welch Road, Palo Alto,
California 94304, USA.
e-mail: ktaymor@stanford.edu
PATENTS
© 2006 Nature Publishing Group http://www.nature.com/naturebiotechnology