© Koninklijke Brill NV, Leiden, 2013 DOI 10.1163/1876312X-44032098 Insect Systematics & Evolution 44 (2013) 117–128 brill.com/ise A new genus and species of Breyeriidae and wings of immature stages from the Upper Carboniferous, Nord-Pas-de-Calais, France (Insecta: Palaeodictyoptera) Jakub Prokop a, *, Zuzana Tippeltová a , Patrick Roques b and André Nel c a Department of Zoology, Faculty of Science, Charles University in Prague Viničná 7, CZ-128 44, Praha 2, Czech Republic b Allée des Myosotis, F-93330 Neuilly sur Marne, France c CNRS UMR 7205, CP 50, Entomologie, Muséum National d’Histoire Naturelle 45 rue Bufon, F-75005 Paris, France *Corresponding author, e-mail: jprokop@natur.cuni.cz Published 10 August 2013 Abstract A new palaeodictyopterid, Aviobreyeria gracilis gen.n., sp.n., is described from an outcrop of Upper Carboniferous (Westphalian C/D) deposits at Avion in the Nord-Pas-de-Calais department (France). his new taxon is placed in the family Breyeriidae based on a comparison of the pattern of venation in its forewing venation with that of other breyeriids and possibly related genera within the Breyerioidea. In addition, wings of diferent immature instars of palaeodictyopterids from the same strata with a similar pattern of venation but difering in having an enlarged RP and a broader anal area (supposed hind wings) were also tentatively attributed to Breyeriidae gen. et sp. indet.. hese wings have a roughly identiiable venation and exhibit the gradual development of wings through diferent ontogenetic stages and their position on the thorax, which is sparsely documented for Paleozoic insects. Discovery of immature and adult stages of palaeodictyopterids in the same layers also indicates they lived in similar habitats. Keywords Late Carboniferous; Moscovian; Palaeodictyopteroida; Breyeroidea; gen.n.; sp.n.; taxonomy; larval wing venation Introduction Breyeriidae is a small family of the Palaeodictyoptera with a characteristic pattern of wing venation and hind wings markedly broader than fore wings, which occur over a rather short interval from the Bashkirian to Gzhelian in Upper Carboniferous deposits in western and central Europe (Belgium, Czech Republic, England, France, Germany and he Netherlands) and USA (Tennessee) (Carpenter 1992; Brauckmann & Gröning 1996; Brauckmann & Herd 2000). Handlirsch (1906–1908) established this family