Growth of Specific Muscle Strength Between 6 and 18
Years in Contrasting Socioeconomic Conditions
Maciej Henneberg,
1
* Gerry Brush,
2
and Geoffrey A. Harrison
2
1
Department of Anatomical Sciences, University of Adelaide, Adelaide, 5005 South Australia, Australia
2
Institute of Biological Anthropology, University of Oxford, Oxford 2OX 6QS, UK
KEY WORDS “Cape Coloured” males and females; Africa; skinfold; arm circumference;
nutrition
ABSTRACT The influence of sex, age, and socioeco-
nomic conditions on specific grip strength of 6 –18-year-old
individuals was studied among 1,704 males and 1,956
females belonging to the so-called “Cape Coloured” com-
munity in the western part of South Africa. Half of the
participants of both sexes came from communities in the
Greater Cape Town area where living conditions are com-
parable to those of middle-class First World communities
(high SES). The other half came from the poorest rural
communities of Klein Karoo (low SES). Arm circumfer-
ences, triceps skinfold thickness, and grip strength of the
right and of the left hand were greater in individuals from
high SES at all ages. Females within each SES group had
skinfolds thicker than males, especially at older ages, and
were weaker. Specific grip strength (SS), estimated as grip
strength per unit area of cross section of the fat-free arm,
increased with age in each group, was greater in males,
and was significantly lower in low SES groups, than in the
high SES ones, especially during and after puberty. It
seems that SES difference in SS will persist into adult-
hood. Sexual differences in SS can be attributed to hor-
monal differences; while the SS increase with age and the
difference between SES groups find no clear explanation
in current theories of muscle growth and development.
Since the speed of neuromuscular reaction observed in our
participants is slower among low SES individuals, it
seems that the difference in neuromuscular control of
strength may be responsible for our findings. Differences
in muscle metabolism and hormonal regulation must also
be considered. Am J Phys Anthropol 115:62–70, 2001.
© 2001 Wiley-Liss, Inc.
It is generally believed that the contractile prop-
erties of human skeletal muscles become mature
early in infancy (Malina and Bouchard, 1991), and
that changes in muscle strength are a direct result
of muscle hypertrophy (Lieber, 1992). Thus, changes
in isometric force as measured by grip strength dur-
ing childhood are attributed mainly to increases in
muscle mass, with some role played by the changing
quality of muscle tissue and/or maturation of neural
control of volitional exertion of maximum force (Ma-
lina, 1986). The growth of muscle fibers, directed by
their use and by maturation of the nervous system,
is still poorly understood (Vogler and Bove, 1985).
Data on changes in muscle composition during
childhood and adolescence are scarce. They indicate
small, if any, changes in the relative contribution of
type I and type II fibers, and an increase in the
amount of intracellular protein and DNA. Due to
small sample sizes and sampling problems, how-
ever, these data should be treated with caution (Ma-
lina, 1986). Besides, it seems that the slow- and
fast-twitch fibers generate similar specific tensions
(Fitts et al., 1991), and thus changes in the propor-
tions of various fiber types during growth should not
affect strength. Although muscle architecture, espe-
cially the angle of fibers to the tendon, affects power
output (Morris, 1948), there seems to be little
change in the basic morphology of muscles during
later childhood and adolescence.
Since the size of muscle fibers seems to be the
major factor responsible for the development of
strength, differences in strength between children
growing in different socioeconomic conditions have
typically been attributed to the differential growth
of muscle mass (Benefice, 1990; Malina and Bou-
chard, 1991). However, it is possible that other fea-
tures of muscle morphology and physiology also play
a significant role (Malina et al., 1987). Particularly
important would be variation in muscle quality as
evidenced, for example, in specific strength, i.e.,
force exerted by a unit of muscle size.
Since strength is one of the basic determinants of
human physical performance, and thus the capacity
for manual labor, it is important to ascertain
Grant sponsor: Medical Research Council (South Africa); Grant
sponsor: Foundation for Research Development; Grant sponsor: Uni-
versity of Cape Town; Grant sponsor: University of Witwatersrand.
*Correspondence to: M. Henneberg, Department of Anatomical Sci-
ences, University of Adelaide Medical School, Adelaide, South Aus-
tralia 5005, Australia. E-mail: maciej.henneberg@adelaide.edu.au
Received 8 April 1999; accepted 31 January 2001.
AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY 115:62–70 (2001)
© 2001 WILEY-LISS, INC.