Behavioural Brain Research 111 (2000) 153 – 163
Research report
Unconditioned anxiety and social behaviour in two rat lines
selectively bred for high and low anxiety-related behaviour
Markus S.H. Henniger * , Frauke Ohl, Sabine M. Ho ¨ lter, Petra Weißenbacher,
Nicola Toschi, Patrick Lo ¨ rscher, Alexandra Wigger, Rainer Spanagel, Rainer Landgraf
Max Planck Institute of Psychiatry, Kraepelinstr. 2 -10, 80804 Munich, Germany
Received 25 October 1999; received in revised form 13 January 2000; accepted 13 January 2000
Abstract
Individuals of high anxiety-related behaviour (HAB) and low anxiety-related behaviour (LAB) rat lines were selectively bred for
differences in anxiety-related behaviour on the elevated plus-maze. The goal of this study was to investigate whether this
behavioural difference is restricted to the test used as the selection criterion or whether it is a stable and robust trait also in other
conflict or non-conflict situations. Therefore, behaviour of male and female HAB and LAB rats was examined in two further tests
of unconditioned anxiety: the black – white box and the social interaction test. Furthermore, behaviour of group-housed male
HAB and LAB rats was studied in their home cages. In addition to standard statistics, discriminant analyses were performed. The
difference in anxiety-related behaviour between the two lines was highly consistent in all tests of unconditioned anxiety. There
were also differences in home cage behaviour, LAB rats being more active than HAB rats; this is likely to be a consequence of
the LAB rats displaying a higher aggressiveness in social behaviour, compared to HAB rats. In all tests used HAB and LAB rats
were clearly distinguished by discriminant analysis. However, while in the elevated plus-maze and the black – white box test the
most important parameters for discrimination between the two lines were mainly those generally seen as closely related to anxiety,
the discrimination in the social interaction paradigm was primarily due to differences in locomotor activity. © 2000 Elsevier
Science B.V. All rights reserved.
Keywords: Emotionality; Elevated plus-maze; Black – white box; Social interaction test; Home cage behaviour
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1. Introduction
We recently reported on two Wistar rat lines which
were selectively bred over several years for extremes in
anxiety-related behaviour on the elevated plus-maze
[12 – 14]: the high anxiety-related behaviour (HAB) and
the low anxiety-related behaviour (LAB) lines. These
lines represent a useful animal model for studying the
neurobiology of inborn anxiety and the mechanism(s)
of actions of anxiolytic compounds [13].
Anxiety has been shown to be a highly complex
construct and it has been suggested that different tests
of anxiety-related behaviour measure various aspects of
anxiety [2,24]. When using animal models of inborn
anxiety it is, therefore, of great interest to evaluate
whether differences in anxiety-related behaviour occur
globally or are restricted to variables measured by the
test used as selection criterion. Therefore, we tested
HAB/LAB rats in different paradigms, which have been
shown to measure distinct dimensions of emotionality.
First, the elevated plus-maze was used to confirm the
selected line differences. Then we tested HAB and LAB
rats in two other tests of unconditioned anxiety, the
black – white box and the social interaction test.
The elevated plus-maze test is a widely used test
paradigm to investigate anxiety-related behaviour (for
reviews see Refs. [6,10,25]) and has extensively been
validated for use in both rats [19] and mice [15]. It is
based on the test-induced conflict in rodents between
* Corresponding author. Tel.: +49-89-30622641; fax: +49-89-
30622569.
E-mail address: henniger@mpipsykl.mpg.de (M.S.H. Henniger)
0166-4328/00/$ - see front matter © 2000 Elsevier Science B.V. All rights reserved.
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