ISSN 1062-8738, Bulletin of the Russian Academy of Sciences. Physics, 2012, Vol. 76, No. 12, pp. 1361–1364. © Allerton Press, Inc., 2012.
Original Russian Text © V.V. Grubov, E.Yu. Sitnikova, A.A. Koronovskii, A.N. Pavlov, A.E. Hramov, 2012, published in Izvestiya Rossiiskoi Akademii Nauk. Seriya Fizicheskaya,
2012, Vol. 76, No. 12, pp. 1520–1523.
1361
INTRODUCTION
There are a great number of effective radiophysical
methods for analyzing and diagnosing the behavior of
complex oscillatory systems. These are widely used in
different fields of natural science in studying and diag-
nosing many systems, including those found in medi-
cine and physiology [1–3]. The application of these
methods in analyzing the rhythmic activity of the
brain is particularly relevant. This activity is a result of
the synchronized operation of a large number of neu-
rons that make up the complex oscillating network of
the brain [1].
Electroencephalograms (EEGs) are traditionally
used in neurophysiological studies to analyze brain
activities [4]. An EEG is the average sum of the cur-
rents generated by the group of neurons in the area of
the recording electrode. Several frequency bands
(alpha, beta, gamma, etc.) are can usually be distin-
guished in an EEG signal. It has been proven that there
is a clear correlation between the nature of the rhyth-
mic activity in an EEG in a specific frequency range
(due to the presence of a rhythm or oscillatory pattern)
and the functional state of the body [1, 4]. An impor-
tant aspect of investigating the nervous system is thus
the study of certain oscillatory patterns, along with the
regularities of their appearance in EEGs in different
states of the organism.
One type of oscillatory activity in an EEG, which
manifests itself during sleep, is sleep spindles, i.e.,
short (0.5–1.5 s) episodes of oscillations with frequen-
cies of 10–16 Hz and a characteristic spindle shape
[5]. It is known that sleep spindles are formed due to
the synchronous operation of the neural network that
consists of the cortex and the thalamus neurons.
The interest in studying sleep spindles is due to
their possible connection with epilepsy. [6] It is known
that the neural network that normally generates sleep
spindles can, under certain conditions, produce sei-
zure activity (i.e., spike wave discharges) [7]. Spike
wave discharges serve as a diagnostic sign of absence
epilepsy, and their presence in an EEG is accompa-
nied by characteristic clinical manifestations. There is
a relationship between the neurophysiological mecha-
nisms of spike-wave discharges and sleep spindles, but
it is complex and far from obvious.
The aim of this work was to study the time-fre-
quency dynamics of sleep spindles (oscillatory pat-
terns) in the EEGs of rats with a genetic predisposition
to absence epilepsy (the WAG/Rij line). Both tradi-
tional wavelet analysis [8] and a new method for
decomposing signals into empirical modes (the Hil-
bert–Huang transform) [9, 10] were used in our anal-
ysis of EEGs.
EXPERIMENTAL DATA
AND TIME-FREQUENCY EEG ANALYSIS
We used EEG records for the cortex and thalamus
of six adult WAG/Rij rats. Recording was continuous
over 24 hours and thus contains fragments of sleep
with strong sleepy spindles and fragments of wakeful-
ness. EEG signals were prefiltered in the range of 0.5–
100 Hz. Our experimental work was performed at the
Institute of Higher Nervous Activity and Neurophysi-
ology of the Russian Academy of Sciences.
Continuous wavelet transform (CWT) [8] was used
for the initial study of EEG signals. As applied to our
Automatic Extraction and Analysis of Oscillatory Patterns
on Nonstationary EEG Signals by Means of Wavelet Transform
and the Empirical Modes Method
V. V. Grubov
a, c
, E. Yu. Sitnikova
b
, A. A. Koronovskii
a, c
, A. N. Pavlov
a
, and A. E. Hramov
a, c
a
Saratov State University, Saratov, 410012 Russia
b
Institute of Higher Nervous Activity and Neurophysiology, Russian Academy of Sciences, Moscow, 117485 Russia
c
Gagarin Saratov State Technical University, Saratov, 410054 Russia
e-mail: vvgrubov@gmail.com
Abstract—The time-frequency structure and dynamics of oscillatory patterns in electroencephalograms of
rats is studied by means of continuous wavelet transform and the decomposition of the signal by empirical
modes. A method for the automatic selection of patterns using the empirical modes is developed. The method
is applied to the study of sleep spindles, and it is shown that their dynamics depends on the regularities of on–
off intermittency.
DOI: 10.3103/S1062873812120167