Cardiovascular Drugs and Therapy 1996;10:251-262.
© Kluwer Academic Publishers, Boston. Printed in U.S.A.
Drug Withdrawal and Rebound Hypertension:
Differential Action of the Central Antihypertensive
Drugs Moxonidine and Clonidine
Heinz Rupp, Bernhard Maisch, and
Christian G. Brilla
Institute of Physiology, University of Tiibingen and Center of
Internal Medicine, Division of Cardiology, Molecular Cardiology
Laboratory, University of Marburg, Marburg, Germany
Summary. To examine the antihypertensive action of the cen-
trally acting antiadrenergic drugs moxonidine and clonidine,
systolic and diastolic blood pressure as well as heart rate
were monitored by radio telemetry in spontaneously hyper-
tensive rats (SHR) with established high blood pressure. In-
creasing doses were administered with regular rat chow for
6-8 day periods. Moxonidine reduced (p < 0.05) diastolic
blood pressure at a dose of 8 mg/kg/day and systolic blood
pressure at 13 mg/kg/day. Heart rate was reduced during high
activity of rats corresponding to an antitachycardiac action.
After withdrawal of 18 mg/kg administered for only 1 day,
blood pressure returned to pretreatment values within 8 days.
Clonidine reduced systolic and diastolic blood pressure at 0.3
mg/kg/day. At 0.8 and 1.3 mg/kg/day, systolic blood pressure
reduction was less pronounced, although heart rate was re-
duced further, reaching values that were below those of un-
treated sleeping rats. When 1.3 mg/kg/day clonidine was dis-
continued, systolic as well as diastolic blood pressure
increased above pretreatment values within I day. A rebound
was also observed in heart rate, which increased by 150 beats/
min. A comparable rebound in blood pressure was observed
after withdrawal of 0.3 mg/kg/day. Since a blood pressure
rebound occured also after withdrawal of 0.3 mg/kg/day clon-
idine in normotensive rats, the rebound phenomenon was in-
dependent of the presence of high blood pressure. No blood
pressure rebound was observed when moxonidine (8 mg/kg/
day) was administered (chow or gavage) in normotensive
rats. These findings in unanesthetized undisturbed rats dem-
onstrate distinct differences in the mode of action of moxo-
nidine and clonidine, which can be accounted for by specific
interactions of moxonidine with imidazoline Ii-receptors,
whereas clonidine would interact not only with Ii-receptors
but also with alpha2-adrenoceptors, and most probably also
with the vagal activity. In view of our previous studies dem-
onstrating a rise in blood pressure and heart rate after a
hypercaloric dietary intake, the selective Ii-receptor agonist
moxonidine appears particularly appropriate for treating
overweight hypertension associated with an enhanced sympa-
thetic outflow of the brain. Of importance in this respect is
that a moxonidine-induced reduction in sympathetic outflow
was not associated with a gain in body weight but resulted
in reduced caloric intake.
Cardiovasc Drugs Ther 1996;10:251-262
Key Words. rebound, hypertension, moxonidine, clonidine,
imidazoline 11receptor, alpha2 receptor
A crucial determinant of primary hypertension is the
sympathetic outflow of the brain. In Western industri-
alized societies, a number of lifestyle factors have
been identified with a synergistic action on sympa-
thetic outflow. Of particular relevance proved to be
nutritional disorders associated with an excess caloric
intake and psychosocial stress [1]. Since an excess
sympathetic outflow of the brain has multiple unfavor-
able actions, drugs that specifically reduce an excess
sympathetic outflow could be efficient in the preven-
tion of catecholamine-mediated organ damage [2-4].
However, until recently centrally acting antihyper-
tensive drugs that reduce sympathetic outflow
showed side effects that limited their general use [5].
Thus, clonidine, which acts on alpha2-adrenoceptors
as well as imidazoline receptors, results in side effects
that are mainly linked to central and peripheral
alphae-adrenoceptors [6-8]. One had thus to rely on
drugs that act peripherally and reduce adrenergic in-
fluences by blocking adrenoceptors. In the last few
years, a second generation of predominantly centrally
acting antihypertensive agents, moxonidine and ril-
menidine, which are selective for imidazoline recep-
tors, became available [9,10]. Moxonidine has a higher
selectivity for imidazoline receptors than rilmenidine
[11]. The imidazoline receptors are located in the ros-
tral ventrolateral medulla of the brain and are in-
volved in excitatory pathways, leading to enhanced
sympathetic outflow. To characterize potential differ-
ences in the action of moxonidine and clonidine, we
used radiotelemetry for monitoring cardiovascular pa-
rameters in conscious spontaneously hypertensive
(SHR) and normotensive Wistar/WU rats.
Address for correspondence:Heinz Rupp, Ph.D., Center of Internal
Medicine, MolecularCardiology Laboratory,Universityof Marburg,
Karl-von-Frisch-Str 1, 35033Marburg, Germany.
Received 7 December1995; accepted4 February 1996
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