African Journal of Biotechnology Vol. 10(59), pp. 12691-12696, 3 October, 2011
Available online at http://www.academicjournals.org/AJB
DOI: 10.5897/AJB10.2359
ISSN 1684–5315 © 2011 Academic Journals
Full Length Research Paper
Intracellular expression of human calcitonin (hCT) gene
in the methylotrophic yeast, Pichia pastoris
Ali Salehzadeh
1
*, Hamideh Ofoghi
2
, Farzin Roohvand
3
, Mohammad Reza Aghasadeghi
3
and Kazem Parivar
1
1
Department of Biology, Faculty of Science, Islamic Azad University, Science and Research Branch, Tehran, Iran.
2
Iranian Research Organization for Science and Technology, Tehran, Iran.
3
Hepatitis and AIDS Department, Pasteur Institute of Iran, Tehran, Iran.
Accepted 20 May, 2011
This study utilized the Pichia pastoris expression system for expression of the synthetic human
calcitonin (hCT) gene, a small peptide hormone secreted by the thyroid gland in mammals and
ultimobranchial glands in lower vertebrate. The P. pastoris vector (pPICZB) contains the alcohol
oxidase gene promoter (AOX1), which under the induction of methanol allows for the expression of
heterologous protein gene inserted downstream in the vector. KM71H (mut
s
) strain of P. pastoris was
used as the host cell. Molecular analysis, including polymerase chain reaction (PCR), sequencing,
restriction enzyme analysis and survival of P. pastoris to increase concentration of zeocin antibiotic
showed that human calcitonin gene was successfully integrated into the P. pastoris genome. The
expected peptide which had an apparent molecular mass of 5.5 kDa was detected by Tricine-SDS-PAGE
analysis and confirmed by enzyme-linked immunosorbent assay (ELISA).
Key words: Pichia pastoris, human calcitonin, KM71H (mut
s
), Tricine-SDS-PAGE.
INTRODUCTION
Calcitonin (CT) is a peptide hormone produced by
specialized C-parafollicular cells of the thyroid glands in
mammals or by cells of the ultimobranchial glands in fish
and reptiles. CT plays an important role in regulating
phosphorus and calcium metabolism, decreasing blood
calcium concentrations and inhibiting bone resorption.
Natural CT and synthesized analog are widely used in
clinical practice for the treatment of postmenopausal
osteoporosis, Paget’s disease of bone, bone pain, spinal
stenosis, acute pancreatitis and gastric ulcer (Li et al.,
2009). Following the increase of the proportion of the
elderly people in the world, osteoporosis has become a
*Corresponding author. E- mail: salehzadehmb@yahoo.com.
Tel: +989126932196.
Abbreviations: CT, Calcitonin; hCT, human calcitonin; SDS-
PAGE, sodium dodecyl sulphate poly acrylamide gel
electrophoresis; PMSF, phenylmethylsulfonyl fluoride.
major threat to the public health due to its high morbidity
and mortality (Lim et al., 2004). Low bone mass and
deterioration of bone micro architecture are the major
characteristics of osteoporosis, which results in increased
bone brittleness and thus is associated with an increased
risk of fracture. CT is one of the effective and safe agents
for the treatment of osteoporosis (Munoz-Torres et al.,
2004). Gills of salmon and pig thyroid glands are the
main source of CT that is used in clinical practice (Tanko
et al., 2004). However, these heterologous products are
short of resources and thus expensive. CT activity is not
species-specific which make it possible to use animal CT
(porcine, salmon and eel) for treatment of human
patients. However, due to immunological reactions, the
prolonged application of animal CT leads to a gradual
decrease or loss of activity. That is why the long term
treatment of human patients with CT requires
homologous human calcitonin (hCT)(Azria 1989). Thus,
genetic engineering techniques with hCT gene as the
target gene may provide solutions to the earlier-
mentioned problem. In this study, we described the