Plant Cell, Tissue and Organ Culture 67: 267–270, 2001.
© 2001 Kluwer Academic Publishers. Printed in the Netherlands.
267
In vitro zygotic embryo culture of wild Musa acuminata ssp. malaccensis
and factors affecting germination and seedling growth
MJ Asif, C Mak & RY Othman
Institute of Biological Sciences, Division of Genetics and Molecular Biology, University of Malaya, 50603 Kuala
Lumpur, Malaysia
Received 1 August 2000; accepted in revised form 30 May 2001
Key words: banana, embryo culture, Musa
Abstract
In vitro zygotic embryo culture of wild banana significantly increased the germination compared to greenhouse
grown seeds. Embryo orientation and BAP concentration significantly affected germination rate. These factors
together with gelling agent, dark and light conditions and coconut water, also showed variable effects on the
number of roots per plant, root length, shoot length, number of days to root emergence and number of days to
shoot emergence.
Introduction
Banana and plantain cultivars are attacked by several
pathogens throughout the world tropics. By far the
most serious pathogen is Fusarium oxysporum f. sp.
cubense. There is no known chemical or cultural con-
trol for the pathogen. Hence, varieties attacked by
this pathogen must be replaced by resistant varieties
(Rowe, 1990). Malaysia is a center of diversity for
both cultivated and wild bananas. The wild progen-
itors of the cultivated bananas co-evolved with the
Fusarium oxysporum f. sp. cubense (Vakili, 1965) of-
fering a possible source of resistance to the disease.
The wild progenitors of the edible bananas produce
seeds while most of the edible clones are seedless with
a few exceptions such as ‘Pisang awak’ subgroup ABB
(Simmonds, 1966). Intractable fertilization barriers
such as moderate to high levels of female sterility and
triploidy make genetic improvement of parthenocarpic
Musa clones slow and technically difficult. Hybrid
plant production in the most common triploid clones
is further complicated by low seed set and germination
(Shepherd et al., 1987). This is due to seed mor-
tality caused by endosperm failure. Embryo culture
before embryo abortion could rescue these interesting
crosses.
Little is known about the factors that affect seed
germination in Musa, except that germination is ex-
tremely variable and relatively difficult to obtain un-
der natural conditions. The use of banana seedlings
as research tools and the increased emphasis on ba-
nana breeding programs requires improved germin-
ation rates. In the current studies, seeds of Musa
acuminata subspecies malaccensis have been used
to optimize in vitro culture of zygotic embryos and
to identify factors affecting germination and seedling
growth.
Materials and methods
Mature seed samples of a diploid Musa acuminata ssp.
malaccensis produced by open pollination were used.
The fruits were thoroughly washed with distilled wa-
ter and then soaked in 1.4% (v/v) sodium hypochlorite
solution for 30 min. The fruit skins were removed, and
seeds were removed under axenic conditions. Seeds
were planted in soil mixture (sand + leaf compost) and
maintained in the greenhouse.
For in vitro culture, seeds were soaked in 1.4%
(v/v) sodium hypochlorite solution for 10 min fol-
lowed by a quick wash with 70% (v/v) ethanol.