Experimental Parasitology 120 (2008) 255–260
0014-4894/$ - see front matter © 2008 Elsevier Inc. All rights reserved.
doi:10.1016/j.exppara.2008.07.011
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Experimental Parasitology
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1. Introduction
Neospora caninum is an obligate intracellular protozoan that
causes abortion and economic loss in the cattle industry. There
are still many aspects of the lifecycle of N. caninum that remain
unknown. At present only the dog and coyote are proven definitive
hosts (McAllister et al., 1998; Gondim et al., 2004) but the recent
finding of oocysts in the faeces of the red fox suggests this may
also be a definitive host (Wapenaar et al., 2006). There is currently
much speculation about the role wildlife may play as a reservoir of
infection (e.g. Gondim, 2006). Recent findings indicate that small
mammals such as mice (Mus domesticus and Mus musculus), rats
(Rattus norvegicus) and wood mice (Apodemus sylvaticus) may be
natural intermediate hosts (Huang et al., 2004; Hughes et al., 2006;
Jenkins et al., 2007; Ferroglio et al., 2007; Barratt et al., 2008). Inves-
tigation of risk factors associated with bovine neosporosis in Nor-
mandy (France) found the presence of rabbits and/or ducks were
a risk factor for seropositivity in dairy cattle (Ould-Amrouche et al.,
1999). To our knowledge there has been no investigation of the
prevalence of N. caninum in wild rabbits in the UK, and only one
previous serological study in rabbits in Spain (Almería et al., 2007).
In this latter study no evidence of exposure to the parasite was
detected in rabbits. The closely related parasite Toxoplasma gondii
has been more extensively studied and has been found to be a natu-
rally occurring infection of wild rabbits in Europe (e.g. Hejlicek and
Literak, 1994; Almería et al., 2004; Figueroa-Castillo et al., 2006).
The objectives of this study were to investigate the prevalence and
co-infection of N. caninum and T. gondii, using PCR techniques,
in a naturally infected population of wild rabbits from Northern
England; the tissue distribution of N. caninum in rabbits and the
range of strain types of T. gondii present in this sample of rabbits.
2. Materials and methods
2.1. Collection of samples and DNA extraction
Rabbits (Oryctolagus cuniculus) n = 57 were collected from
within 2 km of Malham Tarn in the Yorkshire Dales over a three
Neospora caninum: Detection in wild rabbits and investigation of co-infection
with Toxoplasma gondii by PCR analysis
J.M. Hughes
a
, D. Thomasson
a
, P.S. Craig
a
, S. Georgin
a
, A. Pickles
b
, G. Hide
a,
*
a
Centre for Parasitology and Infectious diseases, Biomedical Sciences Research Institute, University of Salford, Salford M5 4WT, UK
b
Field Studies Council at Malham Tarn, Settle, North Yorkshire BD24 9PU, UK
article info abstract
Article history:
Received 9 May 2008
Received in revised form 22 July 2008
Accepted 23 July 2008
Available online 30 July 2008
Neospora caninum is an important pathogen of cattle causing significant economic loss. There is much cur-
rent interest in wild animal reservoirs for this parasite. The role of the rabbit in this is currently unknown.
DNA samples from the brains of wild rabbits (Oryctolagus cuniculus) collected from the Malham area
of the Yorkshire dales were investigated by species-specific PCR for the presence of N. caninum and
Toxoplasma gondii. We found prevalences of N. caninum of 10.5% (6/57) and T. gondii of 68.4% (39/57) with
8.8% (5/57) co-infected. Strain typing of T. gondii positive rabbits revealed strain types I–III were pres-
ent in this population. Investigation of tissue distribution determined N. caninum DNA was most often
detected in the brain and heart, less often in the tongue and not in the liver. To our knowledge this is the
first report of N. caninum detection in naturally infected wild rabbits.
© 2008 Elsevier Inc. All rights reserved.
Keywords:
Apicomlexa
Neospora caninum
Toxoplasma gondii
PCR
Co-infection
Rabbit
Wildlife
DNA, deoxyribonucleic acid
ELISA, enzyme linked immunosorbent
assay
IFAT, indirect florescent test
PCR, polymerase chain reaction
SAG, surface antigen gene
TE, Tris–EDTA buffer
* Corresponding author. Fax: +44 (0)161 2955015.
E-mail address: g.hide@salford.ac.uk (G. Hide).