548 Journal of Basic Microbiology 2010, 50, 548 – 556
© 2010 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim www.jbm-journal.com
Research Paper
Optimization of laccase production by Pleurotus ostreatus
IMI 395545 using the Taguchi DOE methodology
Rathinasamy Periasamy and Thayumanavan Palvannan
Department of Biochemistry, Periyar University, Salem, Tamil Nadu, India
Production of laccase using a submerged culture of Pleurotus orstreatus IMI 395545 was opti-
mized by the Taguchi orthogonal array (OA) design of experiments (DOE) methodology. This
approach facilitates the study of the interactions of a large number of variables spanned by
factors and their settings, with a small number of experiments, leading to considerable savings
in time and cost for process optimization. This methodology optimizes the number of impact
factors and enables to calculate their interaction in the production of industrial enzymes. Eight
factors, viz. glucose, yeast extract, malt extract, inoculum, mineral solution, inducer (1 mM
CuSO
4
) and amino acid (L-asparagine) at three levels and pH at two levels, with an OA layout of
L18 (2
1
× 3
7
) were selected for the proposed experimental design. The laccase yield obtained
from the 18 sets of fermentation experiments performed with the selected factors and levels
was further processed with Qualitek-4 software. The optimized conditions shared an enhanced
laccase expression of 86.8% (from 485.0 to 906.3 U). The combination of factors was further
validated for laccase production and reactive blue 221 decolorization. The results revealed an
enhanced laccase yield of 32.6% and dye decolorization up to 84.6%. This methodology allows
the complete evaluation of main and interaction factors.
Keywords: Laccase / Taguchi DOE methodology / Pleurotus ostreatus IMI 395545 / Optimization /
Dye decolorization
Received: March 14, 2010; accepted: April 29, 2010
DOI 10.1002/jobm.201000095
Introduction
*
Laccase (benzenediol:oxygen oxidoreductase [EC.1.10.3.2])
belongs to a group of enzymes called blue copper oxi-
dases capable of oxidizing phenols and aromatic amines
by reducing molecular oxygen to water. Laccase is
widespread in nature and has been found in plants,
fungi, bacteria and insects [1]. Laccase plays an impor-
tant role in the global carbon cycle and could help in
degrading a wide range of xenoaromatics such as tex-
tile dyes [2], polychlorinated biphenyls, polycyclic aro-
matic hydrocarbons, pesticides and synthetic polymers
[3, 4]. Extensive studies made on fungal laccase have
proved its potential in the various fields of biotechnol-
ogy and created a great market demand for commercial
Correspondence: Dr. T. Palvannan, Department of Biochemistry, Peri-
yar University, Salem – 636 011, Tamil Nadu, India
E-mail: pal2912@yahoo.com
Phone: +91-427-2345766, -2345520
Fax: +91-427-2345124
applications like wastewater detoxification [5, 6], deter-
gent manufacturing and transformation of antibiotics
and steroids [7]. The wide range of applications of lac-
case in the biotechnological and textile industries cre-
ates the need for large amounts of enzymes at low cost
to meet the market demand.
The main limitation for the extensive industrial ap-
plication of laccase is its high cost. To attain the pro-
duction of a large amount of enzyme at low cost, media
optimization plays a crucial role. The optimization of
fermentation media to generate a balanced proportion
of various nutrients is very important to get optimum
microbial growth and enzyme yield [8]. Laccase expres-
sion by fungi is found to be influenced by the applied
culture conditions, such as the nature of the carbon
source, the concentration of a carbon source, the pH of
the fermentation broth, the presence of inducers, the
presence of lignocellulosic materials and a nitrogen
source [5, 9 – 11]. It is essential to optimize all the cul-
ture conditions and the composition of the production